| TREATMENT FROM THE JEPSON MANUAL (1993) |
previous taxon |
next taxon
Jepson Interchange (more information) |
|
|
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora. |
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY |
Perennial or subshrub from caudex or rhizome, generally ± hairy
Stem often ± leafy on lower half, rarely trailing and leafy throughout
Leaves generally simple, basal or sometimes cauline, generally alternate, generally petioled; veins ± palmate
Inflorescence: panicle, generally ± scapose
Flower generally bisexual, generally radial; hypanthium free to ± fused to ovary; calyx lobes generally 5; petals generally 5, free, generally clawed, generally white; stamens generally 5 or 10; pistils 2 and simple or 1 and compound (chambers 1–2, placentas 2–4, axile or parietal), ovary superior to inferior, sometimes more superior in fruit, styles generally 2
Fruit: 2 follicles or 2–4-valved capsule
Seeds generally many, small
Genera in family: 40 genera, 600 species: especially n temp, arctic, alpine; some cultivated (Bergenia , Darmera , Heuchera , Saxifraga , Tellima , Tolmiea )
Reference: [Soltis 1988 Syst Bot 13:64–72]
Rhizome scaly; bulblets 0
Leaves basal, sometimes a few cauline; blade ovate, base cordate to reniform, lobes and teeth shallow, irregular
Inflorescence generally raceme-like; bracts generally scale-like
Flower radial to ± bilateral; hypanthium partly fused to ovary; calyx lobes equal or not; petals 0 or 5, generally equal; stamens 5, generally equal; pistil 1, ovary > half inferior, chamber 1, placentas 2, parietal
Fruit: capsule
Species in genus: 50 species: North America
Etymology: (J.H. von Heucher, German professor of medicine, 1677–1747)
Reference: [Rosendahl, Butters, & Lakela 1936 Minn Stud Plant Sci 2:1–180]
A very difficult genus, highly variable at many levels and needing much additional research.
| Native |
Leaf: petiole 1–9 cm, glandular to stiff-hairy; blade generally < 3 cm wide, broadly ovate, moderately 3–5-lobed, generally truncate to ± cordate
Inflorescence spike-like, 8.5–55 cm, dense, ± short-glandular
Flower ± radial; part of hypanthium fused to ovary 2–2.5 mm, < to > free part, together with calyx lobes 6–8 mm; calyx lobes cream-white to green, sometimes with pink tips; petals generally 0; stamens < calyx lobes, included; mature styles generally < 1 mm, included
Ecology: Rocky banks, slopes
Elevation: 1000–3000 m.
Bioregional distribution: Modoc Plateau
Distribution outside California: to Washington, Idaho, Nevada
2 other vars. from nw US & B.C., with leaf blades generally > 3 cm, generally strongly cordate, occur with var. a. in se OR and may be discovered in MP: var. cylindrica (petiole densely glandular and stiff-hairy) and var. glabella (Torr. & A. Gray) W.E. Wheelock (petiole glabrous to sparsely glandular).