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AMARANTHACEAE

AMARANTH FAMILY

James Henrickson

Annual to tree
Stem prostrate to erect
Leaves alternate or opposite, simple, generally entire, bract-like upward; stipules 0
Inflorescence: cymes, often arrayed ± in spikes or panicles; bracts 1–3 per flower, generally membranous-scarious, tip often short-pointed or spine-like
Flower generally small, uni- or bisexual, radial; sepals 0–5, fused at base or free, often ± scarious; petals 0; stamens 0–5, opposite sepals (staminodes sometimes alternating), filaments sometimes fused at base; ovary superior, 1-chambered, ovules 1–several, erect or pendent on ± basal stalks, styles 0–3, stigma head-like or 2–3-lobed
Fruit: utricle to circumscissile capsule
Seed lenticular to spheric, hard
Genera in family: ± 65 genera, ± 900 species: tropical, subtropical
Reference: [Robertson 1981 J Arnold Arbor 62:267–313]

GUILLEMINEA

Perennial, prostrate, soft-hairy to tomentose, taprooted
Leaves opposite, unequal at a node; petiole widely winged, clasping; blade entire
Inflorescence: spike, generally in axil of shorter leaf, sessile; bracts unequal, white, glabrous, larger ones falling with fruit
Flower bisexual; calyx bell-shaped, lobes 5, ± = tube, glabrous, scarious, tube densely woolly outside; stamens 5, attached to top of calyx tube, filaments fused at base; staminodia 0; ovary compressed-ovoid, style 1, short, stigma ± 2-lobed
Fruit indehiscent; wall membranous
Seed lenticular to spheric, smooth
Species in genus: ± 5 species: s US to South America
Etymology: (A. Guillemin, French botanist, 1796–1842)
Reference: [Henrickson 1987 Sida 12:307–337]

Introduced

G. densa (Willd.) Moq. var. aggregata Uline & Bray

Plant 2–4 dm wide
Leaves: basal in rosettes, 10–40 mm, oblong-lanceolate, early deciduous; cauline petiole 1–6 mm, blade 2–16 mm, elliptic to round, acute to obtuse, glabrous or sparsely silky above, long-soft-hairy below; cauline leaves smaller, clustered in axils
Inflorescence: clusters of spikes, axillary, < or = 4 mm wide, subtended by and mixed with reduced leaves; bracts 1.5–2.5 mm, glabrous, white-scarious
Flower: calyx 1.5–2.8 mm, lobes reflexed, erect in age, tip acute to rounded, midvein ending before tip; filament tube 0.1–0.2 mm, free filaments 0.1–0.3 mm
Seed ± 0.8 mm, red-brown
Ecology: Disturbed places
Elevation: < 100 m.
Bioregional distribution: s Central Coast, South Coast
Distribution outside California: native Arizona to Texas, S.America
Synonyms: Brayulinea d. (Willd.) Small

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