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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

GNAPHALIUM

CUDWEED, EVERLASTING

G. Ledyard Stebbins

Annual or perennial herb, generally ± woolly or tomentose
Leaves alternate, sessile, entire
Inflorescence: heads disciform, many, small, ± sessile in clusters; involucres ± cylindric to spheric, often bell-shaped when pressed; phyllaries graded in several series, transparent to opaque at tips or scarious ± throughout; receptacle flat, naked
Pistillate flowers many, in several series; corollas very slender, minutely lobed, cream to pale yellow or tip reddish
Disk flowers few; corollas ± cylindric to funnel-shaped, whitish to purplish; anther bases short-tailed; style branches wider at tip, truncate
Fruit < 1 mm, oblong; pappus of many fine bristles, bases sometimes fused
Species in genus: ± 120 species: worldwide
Etymology: (Greek: lock of wool)

Native

G. canescens DC.

Biennial or short-lived perennial herb 20–110 cm, much-branched above, scented or not, gray-tomentose throughout; basal leaf-tufts generally present
Leaves 15–80 mm, linear to (ob)lanceolate, sometimes decurrent
Inflorescence panicle-like; heads many; involucre 4.5–6 mm, ovoid; phyllaries white or pale straw-colored, outer ± tomentose (especially base)
Flowers 25–45; pistillate corollas 2.5–4 mm
Fruit: pappus bristles free
Ecology: Many habitats
Elevation: < 2500 m.
Bioregional distribution: North Coast, Klamath Ranges, Outer North Coast Ranges, High Cascade Range, n&c Sierra Nevada Foothills, High Sierra Nevada, Central Western California (except Inner South Coast Ranges), South Coast, n Channel Islands, San Bernardino Mountains, Peninsular Ranges, Great Basin Floristic Province, Desert Mountains
Distribution outside California: to British Columbia, Wyoming, Colorado, New Mexico, n Mexico

Native

subsp. thermale (E.E. Nelson) Stebbins & D.J. Keil

Plant 20–70 cm, ± scented
Leaves 15–80 mm, linear to ± spoon-shaped (basal tufts); cauline ascending, upper decurrent
Inflorescence: branches short, ascending
Flowers 20–35; pistillate corollas 3–3.5 mm
Ecology: Dry woods, roadsides
Elevation: 1000–2500 m.
Bioregional distribution: Klamath Ranges, High Cascade Range, High Sierra Nevada, San Bernardino Mountains, San Jacinto Mountains, Modoc Plateau, White and Inyo Mountains
Distribution outside California: to British Columbia, Wyoming, Colorado

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