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GERANIACEAE

GERANIUM FAMILY

Annual, perennial herb, or ± woody, generally hairy
Leaves simple to compound, basal and cauline; cauline alternate or opposite, stipules present
Inflorescence: cyme or umbel
Flower bisexual, radial or ± bilateral; sepals 5, free, overlapping in bud; petals 5, free, with nectar glands at base; stamens generally 5 or 10; staminodes scale-like or 0; pistil 5-lobed, chambers 5, placentas axile, styles 5, fused to axis, columnar in fruit, stigmas atop axis 5, free
Fruit: segments 5, dry, 1–2-seeded, separating from each other and then from column; fruit body dehiscent on 1 side or not; part of style persistent atop ovary and separating with it, curved to tightly coiled when dry
Genera in family: 14 genera, ± 750 species: temp, ± tropical. Some cultivated for ornamental, perfume oils
Reference: [Robertson 1972 J Arnold Arbor 53:182–201]
Family description, key to genera by M.S. Taylor.

GERANIUM

CRANESBILL, GERANIUM

Mary Susan Taylor

Annual, perennial herb
Leaves palmately lobed or divided; upper alternate or opposite; blade generally round in outline, base generally cordate, ± hairy
Inflorescence: cyme; flowers (1)2
Flower: sepals awned or not; stamens 10, outer 5 opposite petals, inner 5 alternate petals
Fruit: body dehiscent, generally ovoid, 1–2-seeded, base rounded; style column narrowed at top below free stigmas, forming a beak in fruit; part of style persistent to fruit body glabrous to puberulent on side facing column
Species in genus: 250–300 species: temp, tropical mtns
Etymology: (Greek: crane, from beak-like fruit)
Reference: [Jones & Jones 1943 Rhodora 45:5–26;32–53]
Some ornamental, cultivated for oils. Native per (especially G. californicum, G. richardsonii ) vary regionally, are often difficult to separate, need further study.

Introduced

G. anemonifolium L'Hér.

Perennial
Stem ascending to erect, 5–15 dm, glabrous to sparsely hairy, 5–10 mm wide in lower half
Leaves: lower 15–30 cm; blades 8–20 cm wide; leaflets generally 5, 6–10 cm, subsessile, pinnately lobed to dissected
Flower: pedicel 5–10 mm; sepals 8–14 mm, soft-hairy, awned; petals 10–15 mm, rounded, pink to red-purple
Fruit: body 3–4 mm, with net-like ridges, glabrous; style column 16–20 mm, beak 7–9 mm
Seed finely pitted
Ecology: Open to shaded sites
Elevation: < 200 m.
Bioregional distribution: San Francisco Bay Area
Distribution outside California: native to Madeira Island (off nw Africa)

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