RANUNCULACEAE

BUTTERCUP FAMILY

Dieter H. Wilken, except as specified

Annual, perennial herb, sometimes aquatic
Leaves generally basal and cauline, generally alternate, simple or compound; petioles at base generally flat, sometimes sheathing or stipule-like
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; sepals generally 5, free, early deciduous or withering in fruit, generally green; petals 0–many, free; stamens generally 10–many; pistils 1–many, ovary superior, chamber 1, style 1, generally ± persistent in fruit as beak, ovules 1–many
Fruit: achene, follicle, berry, or utricle-like, 1–many-seeded
Genera in family: ± 60 genera, 1700 species: worldwide, especially n temp, tropical mtns; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Erianthis, Helleborus ), some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus )
Reference: [Duncan & Keener 1991 Phytologia 70:24–27]

DELPHINIUM

LARKSPUR

Michael J. Warnock

Perennial; root generally < 10 cm, ± fibrous or fleshy; buds generally obscure
Stem generally 1, erect, generally unbranched; base generally ± as wide as root, generally firmly attached to root, generally ± reddish or purplish
Leaves simple, basal and cauline, petioled; blades generally palmately lobed, deep lobes generally 3–5, generally < 6 mm wide, generally also lobed; lower leaves generally dry, often 0 in flower; cauline merging into bracts upward
Inflorescence: raceme or somewhat branched, terminal; flowers generally 10–25; pedicels generally ± spreading
Flower bilateral; sepals 5, petal-like, generally spreading, generally ± dark blue, uppermost spurred; petals 4, << sepals, upper 2 with nectar-secreting spurs enclosed in uppermost sepal, lower 2 clawed, with blades generally 4–8 mm, notched, generally ± perpendicular to claws, generally colored like sepals, generally obviously hairy; pistils 3(–5)
Fruit aggregate of 3(–5) erect follicles, generally 2.5–4 X longer than wide
Seed dark brown to black, often appearing white, generally winged when immature, generally without inflated collar; coat cell margins generally straight
Etymology: (Latin: dolphin, from bud shape)
Reference: [Lewis & Epling 1954 Brittonia 8:1–22]
Hybrids common, especially in disturbed places. Root length here includes coarse but not thread-like parts. Most species highly TOXIC, attractive and causing many deaths to cattle, less often to horses, sheep.
Horticultural information: Exc as noted, successful In cultivation only within natural range and habitat. Lowland subsp.: DRY. Upland species: winter chilling required.

Native

D. hansenii (Greene) Greene

HANSEN'S LARKSPUR

Stem 25–180 (generally 40–80) cm; base puberulent to hairy
Leaf hairy, especially on lower surface; lobes 3–18; petioles hairy
Inflorescence: flowers often > 25; pedicels 3–57 mm, generally < 8 mm apart, ± ascending, puberulent
Flower: sepals spreading to forward-pointing, lateral 7–13 mm, spur 6–16 mm; lower petal blades generally hairier on inner lobes
Fruit 8–20 mm, often < 3 X longer than wide
Seed finely prickly
Ecology: Oak woodland or open chaparral
Elevation: 60–3000 m.
Bioregional distribution: Inner North Coast Ranges, Cascade Range Foothills, Sierra Nevada Foothills, c&s High Sierra Nevada, Tehachapi Mountain Area, Great Central Valley, w edge Mojave Desert.

Native

subsp. kernense (Davidson) Ewan

Stem 34–110 cm; base puberulent
Leaves generally basal but dry in flower
Inflorescence: pedicels 7–25 (generally > 10) mm apart
Flower: sepals white to dark blue-purple, lateral 7–13 mm, spur 8–16 mm
Chromosomes: 2n=16,32
Ecology: Open oak woodland, chaparral
Elevation: 800–1900 m.
Bioregional distribution: s Sierra Nevada, Tehachapi Mountain Area, w edge Mojave Desert
Flowering time: Apr–May
Horticultural information: DRN, DRY; DFCLT.