|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)45; stamens 45, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):128. See glossary p. 25 for illustrations of general family characteristics.
Annual to perennial herb (sometimes short-lived, dying after flowering once)
Stem generally erect
Leaves: lower generally tapered or petioled, often wavy-margined, generally pinnately lobed, ± dentate, lobes and teeth spine-tipped, margin generally spiny-ciliate, glabrous to tomentose; upper generally sessile, ± reduced
Inflorescence: heads discoid, 1many; involucre cylindric to spheric; phyllaries many, graduated in several series, outer spine-tipped; receptacle flat, long-bristly
Flowers generally many; corollas ± bilateral, white to red or purple, tube long, slender, lobes linear; anther bases sharply sagittate, tips oblong; style tip with slightly swollen node, appendage (above node) long, cylindric, branches very short
Fruit ovoid, glabrous; scar slightly angled; pappus bristles many, plumose, ± persistent or falling in ring
Species in genus: ± 200 species: North America, Eurasia
Etymology: (Greek: thistle)
Taxa difficult, incompletely differentiated, hybridize.
Biennial (or short-lived perennial herb) 429 dm
Stem generally 1, generally simple below; branches few above, ascending, ± white-cobwebby-tomentose, puberulent
Leaves ± persistently gray-tomentose (both surfaces), lighter below; lower 635 cm, petioled or tapered to spiny-winged base, oblong-elliptic to oblanceolate, ± lobed, lobes generally rigidly spreading, simple or with 24 coarse teeth or 2° lobes, main spines 515 mm; middle and upper generally smaller, narrower, decurrent as spiny wings, uppermost well separated, much reduced, ± bract-like, sometimes barely a cluster of long spines
Inflorescence: heads 1few in open cyme-like clusters (sometimes on short axillary branches); peduncles 2.530 cm, leafy; involucres 22.5 cm, 2.55 cm diam, hemispheric or bell-shaped, ± loosely tomentose, sometimes glabrous; phyllaries linear-lanceolate, entire, sometimes midveins with glandular area, outer and middle spreading to reflexed, spines 415 mm, inner with tips ± erect, flat
Flowers: corollas 1827 mm, white to pale lavender or pink, tube 814 mm, throat 47 mm, lobes 59 mm
Fruit 56 mm, dark brown, ± flattened; pappus 1520 mm
Ecology: Canyons, slopes, roadsides
Elevation: 8002100 m.
Bioregional distribution: eastern Mojave Desert
Distribution outside California: to Colorado, New Mexico
Flowering time: AprMay
Closely related to C. occidentale.