|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)45; stamens 45, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):128. See glossary p. 25 for illustrations of general family characteristics.
Annual to perennial herb (sometimes short-lived, dying after flowering once)
Stem generally erect
Leaves: lower generally tapered or petioled, often wavy-margined, generally pinnately lobed, ± dentate, lobes and teeth spine-tipped, margin generally spiny-ciliate, glabrous to tomentose; upper generally sessile, ± reduced
Inflorescence: heads discoid, 1many; involucre cylindric to spheric; phyllaries many, graduated in several series, outer spine-tipped; receptacle flat, long-bristly
Flowers generally many; corollas ± bilateral, white to red or purple, tube long, slender, lobes linear; anther bases sharply sagittate, tips oblong; style tip with slightly swollen node, appendage (above node) long, cylindric, branches very short
Fruit ovoid, glabrous; scar slightly angled; pappus bristles many, plumose, ± persistent or falling in ring
Species in genus: ± 200 species: North America, Eurasia
Etymology: (Greek: thistle)
Taxa difficult, incompletely differentiated, hybridize.
Perennial 522 dm, thinly to densely tomentose, glandular
Leaves generally very strongly wavy; lower 17 dm, petioles spiny-lobed or toothed, blades shallowly to deeply 12 X lobed and dentate, main spines 110 mm, abrupt; cauline gradually reduced, well-distributed, ± lobed, upper clasping (bases expanded, ear-like), sometimes short-decurrent, spines often 1012 mm
Inflorescence: heads ± grouped in panicle-like cluster, ± nodding, closely subtended by uppermost leaves; peduncles 07 cm; involucres hemispheric or bell-shaped, green to purple, 1.53 cm, 24.5 cm diam; phyllaries lanceolate to ovate, spreading to reflexed, spines 15 mm
Flowers: corollas 2022 mm, white to pink or lavender
Fruit 45 mm, ± brown; pappus 1215 mm
Ecology: Serpentine seeps and streams
Elevation: < 750 m.
Bioregional distribution: San Francisco Bay Area, c Outer South Coast Ranges.
See the CNPS Inventory for information about endangerment and rarity.
Stem < 2 m, green to purplish
Leaves densely tomentose, especially lower surface; glandular hairs concealed
Inflorescence: heads ± nodding in flower, generally erect in fruit; outer phyllaries 1520 mm, green to dark purple, strongly recurved, ± channeled, widest below middle, upper 1/2 tapered to spine 14 mm
Flowers: corolla tube 7 mm, throat 8 mm, lobes 5 mm; style branches 46 mm
Fruit minutely scabrous above
Ecology: Habitat of sp.
Elevation: < 300 m.
Bioregional distribution: c Outer South Coast Ranges (San Luis Obispo Co.).