Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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RANUNCULACEAE

BUTTERCUP FAMILY

Dieter H. Wilken, except as specified

Annual, perennial herb, sometimes aquatic
Leaves generally basal and cauline, generally alternate, simple or compound; petioles at base generally flat, sometimes sheathing or stipule-like
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; sepals generally 5, free, early deciduous or withering in fruit, generally green; petals 0–many, free; stamens generally 10–many; pistils 1–many, ovary superior, chamber 1, style 1, generally ± persistent in fruit as beak, ovules 1–many
Fruit: achene, follicle, berry, or utricle-like, 1–many-seeded
Genera in family: ± 60 genera, 1700 species: worldwide, especially n temp, tropical mtns; many ornamental (Adonis, Aquilegia, Clematis, Consolida, Delphinium, Erianthis, Helleborus ), some highly TOXIC (Aconitum, Actaea, Delphinium, Ranunculus )
Reference: [Duncan & Keener 1991 Phytologia 70:24–27]

AQUILEGIA

COLUMBINE

Perennial from thick, simple to branched caudex
Stems 1–few, ascending to erect, branched to not, scapose to not, glabrous to glandular-hairy
Leaves: basal 1–3-ternate, petiole generally long; cauline 0–few, generally much reduced, deeply 3-lobed to 1–2-ternate, petiole short to ± 0; segments generally wedge-shaped to obovate, upper surface green to pale green, lower surface pale green to glaucous
Inflorescence: few-flowered raceme or flower solitary, terminal; axis and pedicels glabrous to glandular; flowers often pendent
Flower radial; sepals 5, petal-like, spreading to slightly reflexed; petals 5, generally with spur projecting between sepals; pistils generally 5
Fruit: follicles, glabrous to glandular
Seeds smooth, shiny, brown to black
Species in genus: ± 70 species: temp North America, Eurasia
Etymology: (Derivation uncertain, perhaps Latin: eagle, from spurs, or water-drawer, from habitats)
Reference: [Munz 1946 Gentes Herb 7:1–150]
Many species and hybrids cultivated as ornamental; natural hybrids common.

Native

A. formosa DC.

Plant 20–80(150) cm
Leaves: basal and lower cauline 2–3-ternate, petioles 5–30(40) cm, segments 7–45(130) mm; upper cauline generally simple to deeply 3-lobed
Inflorescence: flower nodding, in fruit erect
Flower: sepals 12–20(25) mm, red; petal blade 1–8 mm, yellow, spur (4)10–23 mm, ± straight to ± incurved, tube red, tip 1.5–4 mm wide, mouth 4–8 mm wide, ± round, ± 90° to flower axis, red to yellowish; stamens 10–18 mm
Chromosomes: 2n=14
Ecology: Streambanks, seeps, moist places, chaparral, oak woodland, mixed-evergreen or coniferous forest
Elevation: < 3300 m.
Bioregional distribution: California Floristic Province (except Great Central Valley, South Coast, Channel Islands), Great Basin Floristic Province, Desert Mountains
Distribution outside California: to Alaska, Montana, Baja California
Flowering time: Jul
Synonyms: vars. hypolasia (Greene) Munz, pauciflora (Greene) H.S. Boothman, truncata (Fisch. & C.A. Mey.) Baker; forma anomala J.T. Howell
Lf variation needs study; plants with pale green to glaucous, generally 3-ternate leaves (DMtns), have been called A. shockleyi Eastw
Horticultural information: IRR, DRN: 4, 5, 6, 15, 16, 17, 24 &SHD: 1, 2, 3, 7, 8, 9, 14, 18, 19, 20, 21, 22, 23.

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bioregional map for AQUILEGIA%20formosa being generated
 
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Aquilegia formosa
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