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BERBERIDACEAE

BARBERRY FAMILY

Michael P. Williams

Perennial, shrub, generally from rhizomes; caudex sometimes present, glabrous, glaucous, or hairy
Stems spreading to erect, branched or not
Leaves simple, 1–3-ternate, or pinnately compound, basal and cauline, generally alternate, deciduous or evergreen, petioled
Inflorescence: generally raceme, spike, or panicle, scapose, terminal, or axillary
Flower: sepals 6–18 or 0, generally in whorls of 3; petals generally 6, in 2 whorls of 3, or 0; stamens 6–12, free or fused at base, 2-whorled or not, anthers dehiscent by flap-like valves or longitudinal slits; ovary superior, chamber 1, ovules generally 1–10, style 1 or 0, stigma flat or spheric
Fruit: berry, capsule, or achene
Genera in family: 16 genera, ± 670 species: temp, tropical worldwide; some cultivated (Berberis, Epimedium, Nandina (Heavenly bamboo), Vancouveria )
Reference: [Ernst 1964 J Arnold Arbor 45:1–35]

ACHLYS

VANILLA LEAF, DEER FOOT

Perennial from scaly rhizomes; caudex short, erect
Leaves 1–few, basal, long-petioled, ternate; leaflets triangular to fan-shaped, base tapered
Inflorescence: spike, ± scapose, dense, long-peduncled; lateral flowers generally unisexual, terminal flowers bisexual
Flower: perianth 0; stamens generally 9, anther valves flap-like, curled inward; ovule 1, style 0, stigma ± flat, furrowed
Fruit: achene, curved, furrowed, brown to red-purple
Species in genus: 2 species: w North America, Japan
Etymology: (Greek: thin mist or obscurity, from inconspicuous flowers)
Reference: [Terabayashi 1981 Bot Mag Tokyo 94:141–157]

Native

A. californica I. Fukuda & H.G. Baker

Plant 3–5 dm; rhizome internodes (7.5)9–10 cm
Leaf: terminal leaflet generally 7–16 cm, 8–17 cm wide, (3)6–8(12)-lobed
Inflorescence (except peduncle) < 4 cm
Flower: stamens 4–6 mm; ovary 1.5–2 mm
Fruit 3.5–5 mm, brown
Chromosomes: 2n=24
Ecology: Moist, shaded sites, coniferous forest
Elevation: < 1200 m.
Bioregional distribution: North Coast, w Klamath Ranges, w Outer North Coast Ranges
Distribution outside California: to British Columbia
Horticultural information: TRY.

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