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PTERIDACEAE

BRAKE FAMILY

Alan R. Smith and Thomas Lemieux

Perennial, in soil or on or among rocks; rhizome creeping to erect, scaly
Leaves generally all ± alike (or of 2 kinds, fertile and sterile), generally < 50 cm, often < 25 cm; petiole generally thin, wiry, often dark, in transverse section with vascular strands generally 1–3, less often many in a circle; blade generally pinnate or ± palmate-pinnate (see Adiantum ), often 2 or more compound, lower surface often with glands, ± powdery exudate, hairs, or scales; segments round, oblong, fan-shaped, or otherwise, veins generally free
Sporangia in sori or not, marginal, submarginal, or along veins, sometimes covered by recurved, often modified segment margins (false indusia); true indusia 0; spores spheric, sides sometimes flat, scar with 3 radiating branches
Genera in family: ± 40 genera, 500 species: worldwide, especially dry areas. Definition of Cheilanthes and related genera problematic; traditional limits often untenable.

ASTROLEPIS

Plant in soil or rock crevices; rhizome ± short-creeping-decumbent, scales generally linear to linear-lanceolate, toothed, pale to reddish brown, older with a darker, irregular central area or not
Leaf: axes generally orange to reddish brown, scaly; blade 1-pinnate, linear, leaflets sometimes shallowly but not deeply pinnately lobed or dissected, upper surface with stellate scales
Sporangia along veins, obscured by dense scales; segment margin unmodified, not recurved
Species in genus: ± 6 species: sw US through South America
Etymology: (Greek: star scale)
Reference: [Benham et al. 1988 Amer J Bot 75(6:2):138]

Native

A. cochisensis (Goodd.) D.M. Benham & Windham

SCALY CLOAK FERN

Rhizome short; scales ± 10 mm, 0.1–0.5 mm wide, linear; teeth ± sparse, more pronounced in upper half or not
Leaf: petiole 3–6(10) cm, 1–1.5 mm wide, scales appressed, 0.5 mm, whitish; blade 1-pinnate, 8–15(20) cm, tapered to tip; 1° leaflets < 0.5 cm, with jointed stalk and 0–3 pairs of lobes, generally obtuse at tip, upper surface with persistent, stellate scales, lower surface with lanceolate, whitish to tan, abundantly toothed and finely dissected scales covering small (< 0.1 mm) glandular hairs
Sporangia in a submarginal band when mature, ± visible, erupting through scales
Ecology: Limestone slopes, crevices
Elevation: 900–1800 m.
Bioregional distribution: Desert Mountains
Distribution outside California: to Texas, Mexico, including Baja California
Synonyms: Cheilanthes c. (Goodd.) Mickel; C. sinuata (Sw.) Domin var. c. (Goodd.) Munz

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