TREATMENT FROM THE JEPSON MANUAL (1993) |
previous taxon |
next taxon
Jepson Interchange (more information) |
|
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora. |
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY |
Perennial, on plants, rocks, in rock crevices, less often in soil, humus, or on dunes; rhizome short- to long-creeping, branched, glaucous to not, scaly
Leaves ± alike or of 2 kinds, fertile and sterile; petiole thin to thick, straw-colored or green to brown or black, generally jointed to persistent knob on rhizome; blade generally simple to 1-pinnate, membranous to fleshy or leathery; veins free to fused, generally forked
Sporangia: sori round to elongate, rarely linear, generally 1 per areole, in 1several rows on each side of segment midrib; indusium 0; spores generally ± elliptic, ± smooth to coarse-tubercled or -ridged, scar linear
Genera in family: ± 46 genera, ± 650 species: worldwide, especially Old World tropical. Numbers of genera, species depend on treatment; many species cultivated.
Rhizome long-creeping; scales lanceolate, ± brownish, 1-colored or often with darker central area or midstripe
Leaves 0.210(20) dm, ± alike or fertile > sterile; petiole glabrous to scaly; blade 1-pinnate to generally deeply pinnately lobed, rarely simple and unlobed, glabrous to hairy, glandular or not, scales on lower surface midrib near base generally lanceolate or linear-lanceolate, generally ± brown; veins free to fused
Sporangia: sori in 1 row on each side of segment midrib, generally raised, sometimes including branched or unbranched, glandular hairs, sporangium-like structures, or shriveled sporangia; spores yellow
Species in genus: ± 160 species: generally New World, tropical, some temp, few boreal
Etymology: (Latin: many feet, from rhizome)
Reference: [Whitmore & Smith 1991 Madroño 38:233248]
50% or more malformed spores indicates hybrid involving 2 or more species in CA.
Native |
Rhizome 312 mm diam, conspicuously whitish glaucous, taste bland; scales with darker midstripe, often partly deciduous in age
Leaves evergreen 1several seasons; blade 618(50) cm, deltate to oblong-ovate, thick, leathery, firm, brittle, upper sides of midribs glabrous, lower with deltate, dark brown to black-brown scales near base, segments crenate, margins like cartilage, tips generally round or round-obtuse, veins ± often fused
Sporangia: sori 25 mm, ovate to round, often ± merged, sometimes with short, branched, glandular hairs
Chromosomes: 2n=74
Ecology: Coast, generally in heavy fog-drip or salt-spray zones, on plants, dunes, rocky cliffs, bluffs, mtn ridges, granitic or volcanic rocks, mossy logs, or in soil
Elevation: 0610 m.
Bioregional distribution: North Coast, Outer North Coast Ranges, Central Coast, San Francisco Bay Area
Distribution outside California: to British Columbia, Baja California (Guadalupe Island)
Plants called P. scouleri from n ChI are fertile or sterile hybrids (2n=111,148) involving P. californicum
Horticultural information: DRN, SHD: 4, 5 &IRR: 6, 16, 17, 24.