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SOLANACEAE

NIGHTSHADE FAMILY

Michael Nee

Annual to shrub
Leaves generally simple, generally alternate, generally petioled; stipules 0; blade entire to deeply lobed
Inflorescence various
Flower bisexual; calyx lobes generally 5; corolla ± radial, cylindric to rotate, lobes generally 5; stamens 5, alternate corolla lobes; ovary superior, generally 2-chambered, style 1
Fruit: berry or capsule, 2–5-chambered
Genera in family: 75 genera, 3000 species: worldwide, especially ± tropical; many alien weeds in CA; many cultivated for food, drugs, or ornamental (potato, tomato, peppers, tobacco, petunia); many TOXIC.

SOLANUM

NIGHTSHADE

Annual to shrub or vine, often glandular, sometimes prickly
Leaves alternate to subopposite, often unequal, entire to deeply pinnately lobed
Inflorescence: panicle or umbel-like, often 1-sided
Flower: calyx ± bell-shaped; corolla ± rotate, white to purple; anthers free, > filaments, oblong or tapered, opening by 2 pores or short slits near tip; ovary 2-chambered, style 1, stigma head-like
Fruit: berry, generally spheric (or dry, capsule-like)
Seeds many, compressed, generally reniform
Species in genus: ± 1500 species: worldwide, especially tropical Am
Etymology: (Latin: quieting, from narcotic properties)
Reference: [Symon 1981 J Adelaide Bot Gard 4:1–367]
Many cultivated for food (including potato, S. tuberosum ), ornamental; many TOXIC.

Introduced

S. marginatum L.

WHITE-MARGINED NIGHTSHADE

Shrub < 2 m, prickly, densely stellate-hairy
Leaf 10–18 cm, widely ovate, wavy-lobed, densely white-tomentose below
Inflorescence raceme-like; lowermost flower bisexual, reflexed, others staminate
Flower: calyx ± 10 mm, prickly; corolla 15–30 mm wide, white; anthers 6–7 mm
Fruit 35–50 mm diam, tough, yellow, glabrous
Seed ± 3 mm
Chromosomes: n=12
Ecology: Disturbed places
Elevation: generally < 1000 m.
Bioregional distribution: Central Coast, South Coast
Distribution outside California: native to Africa
Weedy.

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