Jepson Flora Project: Jepson Interchange    

link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.



Michael Nee

Annual to shrub
Leaves generally simple, generally alternate, generally petioled; stipules 0; blade entire to deeply lobed
Inflorescence various
Flower bisexual; calyx lobes generally 5; corolla ± radial, cylindric to rotate, lobes generally 5; stamens 5, alternate corolla lobes; ovary superior, generally 2-chambered, style 1
Fruit: berry or capsule, 2–5-chambered
Genera in family: 75 genera, 3000 species: worldwide, especially ± tropical; many alien weeds in CA; many cultivated for food, drugs, or ornamental (potato, tomato, peppers, tobacco, petunia); many TOXIC.



Annual or rhizomed per; hairs sometimes branched
Leaves sometimes ± opposite, entire to pinnately lobed
Inflorescence: flowers 1–few per axil, pedicelled
Flower: calyx 5-lobed, enlarged and persistent in fruit; corolla rotate to widely bell-shaped, yellowish, often dark-spotted inside; stamens 5, filaments inserted on hairy band in corolla tube, anthers free, generally < filaments, opening by slits; style generally straight
Fruit: berry
Seeds many, 2–2.5 mm, ± spheric to reniform
Species in genus: ± 85 species: Am, Eurasia, Australia
Etymology: (Greek: bladder, from calyx in fruit)
Reference: [Sullivan 1985 Syst Bot 10:426–444]
Some species cultivated for edible or ornamental fruit. Unripe fruit often TOXIC. Needs further study in w US.


P. hederifolia A. Gray

Perennial or subshrub 1–8 dm, from fleshy rhizome; hairs sometimes branched or glandular
Leaf 2–4 cm, ovate, entire to coarsely toothed, generally gray-green; base tapered or subcordate
Inflorescence: pedicels 3–5(10) mm, in fruit < 15 mm
Flower: calyx 6–7 mm, in fruit 20–30 mm, ± spheric, with 10 green veins; corolla 10–15 mm, widely bell-shaped, yellow, generally with 5 purple-brown spots at base inside
Ecology: Gravelly to rocky slopes
Elevation: 700–1800 m.
Bioregional distribution: s Peninsular Ranges, s East of Sierra Nevada, Mojave Desert
Distribution outside California: to Texas, n Mexico


var. fendleri (A. Gray) Cronquist

Hairs generally branched, nonglandular
Chromosomes: n=12
Ecology: Habitats of sp
Elevation: 900–1800 m.
Bioregional distribution: s East of Sierra Nevada (including Inyo Mtns), Mojave Desert
Distribution outside California: to Texas, n Mexico
Flowering time: May–Jul
Synonyms: var. cordifolia (A. Gray) Waterf.; P. f. A. Gray

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bioregional map for PHYSALIS%20hederifolia%20var.%20fendleri being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Physalis hederifolia var. fendleri
Retrieve dichotomous key for Physalis
Overlay Consortium of California Herbaria specimen data by county on this map
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