Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.

OPHIOGLOSSACEAE

ADDER'S-TONGUE FAMILY

Warren H. Wagner, Jr.

Perennial, small, fleshy, generally glabrous; caudex generally underground, unbranched; roots glabrous, with bulblets or plantlets or not
Leaf generally 1 per caudex per year, divided into 2 facing parts with a common stalk; sterile part separated from fertile at to well above ground, blade simple to compound, veins free and forked (or netted, with included veinlets); fertile part bladeless, bearing sporangia, simple to compound
Sporangia dehiscent into 2 valves, ± 1 mm wide, thick-walled
Genera in family: 3 genera, 70–85 species: ± worldwide, generally rare or overlooked. Fern-like plants with many traits of seed plants. Specimens must be carefully spread and pressed for identification; haploid generation underground, fleshy, non-green, associated with fungi.

BOTRYCHIUM

GRAPE-FERN, MOONWORT

Roots smooth, pale or cork-ridged, dark gray, without bulblets or plantlets
Leaf generally deciduous; bud glabrous or hairy; sterile part generally ± 1–3-pinnate (rarely simple or entire), linear to deltate, segments linear to oblong and midribbed or spoon- to wedge- or fan-shaped and not midribbed, veins free, forked, margins entire to dentate or irregularly cut; fertile part 1–3-pinnate, < to > sterile
Sporangia not sunken in axis; stalk 0 or short
Species in genus: 40–50 species: generally temp to arctic or alpine
Etymology: (Greek: bunch of grapes, from clusters of sporangia)
Reference: [Wagner & Wagner 1983 Amer Fern J 73:53–62]
Difficult, needing careful study; most species very uncommon, sporadic; good sampling of populations highly desirable in specimens
Horticultural information: TRY; DFCLT.

Native

B. pinnatum H. St. John

NORTHWESTERN MOONWORT


Leaf: sterile part separated from fertile near middle, stalk very short to 0, blade deeply lobed to 2-pinnate, < 8 cm, < 5 cm wide, oblong to deltate, segments ± ascending, < 7 pairs, oblique-ovate to lanceolate-oblong, shiny, bright green, midribbed, margins entire to minutely crenate; fertile part 2-pinnate, 1–2 X sterile
Chromosomes: 2n=180
Ecology: Fields, shrubby slopes
Elevation: probably ± 2000 m.
Bioregional distribution: Klamath Ranges (near Etna Mills, Siskiyou Co.), High Cascade Range (Mount Shasta)
Distribution outside California: to Alaska, Colorado
Synonyms: B. boreale subsp. obtusilobatum (Rupr.) R.T. Clausen
B. lanceolatum (C.C. Gmel.) Angström (sterile leaf part triangular, fertile leaf part branched into 2–4 major axes near base) would key here, might be discovered in mtns of n CA. B. pumicola Coville, Oregon moonwort, of Crater Lake region, OR, has not been found in CA.
See the CNPS Inventory for information about endangerment and rarity.
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bioregional map for BOTRYCHIUM%20pinnatum being generated
 
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Botrychium pinnatum
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