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This page is based on the 1993 Jepson Manual.
Please see the Jepson eFlora for up-to-date information about California vascular plants. |
| Jepson Flora Project: Jepson Interchange |
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TREATMENT FROM THE JEPSON MANUAL |
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Jepson Interchange (more information) |
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©Copyright 1993 by the Regents of the University of California
Print edition is available from the University of California Press |
| The second edition of The Jepson Manual (2012) is available from the University of California Press | |
| See also the Jepson eFlora, which parallels the Second Edition |
Perennial, small, fleshy, generally glabrous; caudex generally underground, unbranched; roots glabrous, with bulblets or plantlets or not
Leaf generally 1 per caudex per year, divided into 2 facing parts with a common stalk; sterile part separated from fertile at to well above ground, blade simple to compound, veins free and forked (or netted, with included veinlets); fertile part bladeless, bearing sporangia, simple to compound
Sporangia dehiscent into 2 valves, ± 1 mm wide, thick-walled
Genera in family: 3 genera, 7085 species: ± worldwide, generally rare or overlooked. Fern-like plants with many traits of seed plants. Specimens must be carefully spread and pressed for identification; haploid generation underground, fleshy, non-green, associated with fungi.
Roots smooth, pale or cork-ridged, dark gray, without bulblets or plantlets
Leaf generally deciduous; bud glabrous or hairy; sterile part generally ± 13-pinnate (rarely simple or entire), linear to deltate, segments linear to oblong and midribbed or spoon- to wedge- or fan-shaped and not midribbed, veins free, forked, margins entire to dentate or irregularly cut; fertile part 13-pinnate, < to > sterile
Sporangia not sunken in axis; stalk 0 or short
Species in genus: 4050 species: generally temp to arctic or alpine
Etymology: (Greek: bunch of grapes, from clusters of sporangia)
Reference: [Wagner & Wagner 1983 Amer Fern J 73:5362]
Difficult, needing careful study; most species very uncommon, sporadic; good sampling of populations highly desirable in specimensHorticultural information: TRY; DFCLT.
| Native |
Leaf: sterile part separated from fertile generally above middle of leaf, stalk < 7 mm, blade 1-pinnate, generally < 6 cm, < 2 cm wide, oblong, thin, soft, shiny, yellow-green, segments ± well separated, 35 pairs, not midribbed, lower widely fan-shaped, with margins at base meeting at 120160°, outer margins generally finely crenate to dentate; fertile part 12-pinnate, 1.33 X sterile
Chromosomes: 2n=90
Ecology: Marshes, meadows
Elevation: 15002500 m.
Bioregional distribution: High North Coast Ranges, Cascade Range, Sierra Nevada, San Gabriel Mountains, San Bernardino Mountains
Distribution outside California: to Washington, Utah
Extremely local; often with B. simplex.
| YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps). |
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