Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

OPHIOGLOSSACEAE

ADDER'S-TONGUE FAMILY

Warren H. Wagner, Jr.

Perennial, small, fleshy, generally glabrous; caudex generally underground, unbranched; roots glabrous, with bulblets or plantlets or not
Leaf generally 1 per caudex per year, divided into 2 facing parts with a common stalk; sterile part separated from fertile at to well above ground, blade simple to compound, veins free and forked (or netted, with included veinlets); fertile part bladeless, bearing sporangia, simple to compound
Sporangia dehiscent into 2 valves, ± 1 mm wide, thick-walled
Genera in family: 3 genera, 70–85 species: ± worldwide, generally rare or overlooked. Fern-like plants with many traits of seed plants. Specimens must be carefully spread and pressed for identification; haploid generation underground, fleshy, non-green, associated with fungi.

BOTRYCHIUM

GRAPE-FERN, MOONWORT

Roots smooth, pale or cork-ridged, dark gray, without bulblets or plantlets
Leaf generally deciduous; bud glabrous or hairy; sterile part generally ± 1–3-pinnate (rarely simple or entire), linear to deltate, segments linear to oblong and midribbed or spoon- to wedge- or fan-shaped and not midribbed, veins free, forked, margins entire to dentate or irregularly cut; fertile part 1–3-pinnate, < to > sterile
Sporangia not sunken in axis; stalk 0 or short
Species in genus: 40–50 species: generally temp to arctic or alpine
Etymology: (Greek: bunch of grapes, from clusters of sporangia)
Reference: [Wagner & Wagner 1983 Amer Fern J 73:53–62]
Difficult, needing careful study; most species very uncommon, sporadic; good sampling of populations highly desirable in specimens
Horticultural information: TRY; DFCLT.

Native

B. ascendens W.H. Wagner

UPSWEPT MOONWORT


Leaf: sterile part separated from fertile near or generally above middle of leaf, stalk < 1 cm, blade 1-pinnate, < 6 cm, < 5 cm wide, oblong to oblong-dentate, thin but firm, veiny, yellow-green, segments well separated, ascending, < 5 pairs, obliquely wedge-shaped, not midribbed, lower with margins at base meeting at 40–90(120)°, outer margins sharply and finely dentate to cut, basal segments often with scattered marginal sporangia; fertile part 1–2-pinnate, 1.3–2 X sterile
Chromosomes: 2n=180
Ecology: Grassy fields, coniferous woods near streams
Elevation: 1500–1800 m.
Bioregional distribution: s High Cascade Range (near Jonesville, Butte, Tehama cos.)
Distribution outside California: to British Columbia, Nevada
See the CNPS Inventory for information about endangerment and rarity.
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