| TREATMENT FROM THE JEPSON MANUAL (1993) |
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©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora. |
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY |
Annual to shrubs, generally glandular, some green root-parasites
Stem generally round
Leaves generally alternate, simple, generally ± entire; stipules generally 0
Inflorescence: spike to panicle, generally bracted, or flowers 1–2 in axils
Flower bisexual; calyx lobes generally 5; corolla generally strongly bilateral, generally 2-lipped (upper lip generally 2-lobed, lower lip generally 3-lobed); stamens generally 4 in 2 pairs, generally included, a 5th (generally uppermost) sometimes present as a staminode; pistil 1, ovary superior, chambers generally 2, placentas axile, style 1, stigma lobes generally 2
Fruit: capsule, generally ± ovoid, loculicidal or septicidal
Seed: coat sculpture often characteristic
Genera in family: ± 200 genera, 3000 species: ± worldwide; some cultivated as ornamental (e.g., Antirrhinum, Mimulus, Penstemon ) or medicinal (Digitalis )
Recent taxonomic note: Recently treated to include only Buddleja, Scrophularia, and Verbascum in CA; other genera moved to Orobanchaceae (Castilleja, Cordylanthus, Orthocarpus, Pedicularis, Triphysaria), Phrymaceae (Mimulus), and Plantaginaceae (= Veronicaceae sensu Olmstead et al.)
Key to genera by Elizabeth Chase Neese & Margriet Wetherwax.
Perennial or shrub, long-soft-shaggy- and short-stout-glandular-hairy
Stem square
Leaves opposite, 4-ranked, lanceolate to triangular-ovate, petioled
Inflorescence: panicle; pedicels > bracts; bracts lanceolate, long-tapered
Flower: calyx lobes 5, obtuse to rounded; corolla 5-lobed, 2-lipped, greenish yellow to blackish, tube urn-shaped to spheric, lowest lobe generally recurved; fertile stamens 4, included, anther sacs 1, staminode attached near base of upper lip, appressed to upper side of corolla; style slender, generally reflexed at flower, stigma head-like; nectary at base of ovary, fleshy
Fruit septicidal
Seeds many, oblong-ovoid, ridged
Species in genus: ± 150 species: n temp, especially Eurasia
Etymology: (Latin: scrofula, a disease supposedly cured by some species)
| Native |
Shrub, glandular-puberulent
Stem 12–18 dm
Leaf: blade 10–15 cm, ovate, truncate to cordate, long-tapered, sharply dentate; petiole 3–5 cm
Inflorescence long, narrowly branched, long-soft-wavy-glandular-hairy
Flower: calyx lobes 3 mm, triangular-ovate, acute; corolla 8–11 mm, spheric to urn-shaped, blackish maroon, lowest lobe recurved; staminode 0 or awl-like vestige, maroon
Fruit 6–8 mm, conic
Ecology: Canyons, coastal scrub, chaparral
Elevation: < 500 m.
Bioregional distribution: s Channel Islands (Santa Catalina, San Clemente islands).