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SCROPHULARIACEAE

FIGWORT FAMILY

Lawrence R. Heckard, Family Coordinator

Annual to shrubs, generally glandular, some green root-parasites
Stem generally round
Leaves generally alternate, simple, generally ± entire; stipules generally 0
Inflorescence: spike to panicle, generally bracted, or flowers 1–2 in axils
Flower bisexual; calyx lobes generally 5; corolla generally strongly bilateral, generally 2-lipped (upper lip generally 2-lobed, lower lip generally 3-lobed); stamens generally 4 in 2 pairs, generally included, a 5th (generally uppermost) sometimes present as a staminode; pistil 1, ovary superior, chambers generally 2, placentas axile, style 1, stigma lobes generally 2
Fruit: capsule, generally ± ovoid, loculicidal or septicidal
Seed: coat sculpture often characteristic
Genera in family: ± 200 genera, 3000 species: ± worldwide; some cultivated as ornamental (e.g., Antirrhinum, Mimulus, Penstemon ) or medicinal (Digitalis )
Recent taxonomic note: Recently treated to include only Buddleja, Scrophularia, and Verbascum in CA; other genera moved to Orobanchaceae (Castilleja, Cordylanthus, Orthocarpus, Pedicularis, Triphysaria), Phrymaceae (Mimulus), and Plantaginaceae (= Veronicaceae sensu Olmstead et al.)
Key to genera by Elizabeth Chase Neese & Margriet Wetherwax.

PENSTEMON

BEARDTONGUE

Noel H. Holmgren

Perennial to shrub
Leaves generally opposite, entire to toothed; upper sessile
Inflorescence: panicle or raceme; bracts generally small
Flower: calyx lobes 5, ± equal; corolla tube ± cylindric or lower side expanded, ± 2-lipped, generally pink or blue to purple (some red, yellow, or white), upper lip 2-lobed, external in bud; anther sacs 2, generally spreading ± flat at dehiscence; staminode attached near base of corolla tube, well developed, generally hairy on upper side; nectaries 2, at bases of upper stamens; stigma head-like
Fruit: capsule, septicidal and sometimes also loculicidal at tip
Seeds generally many, irregularly angled
Species in genus: 250 species: North America., especially w US
Etymology: (Latin & Greek: almost thread, from stamen-like staminode)
Reference: [Holmgren 1984 In Cronquist et al. Intermountain Flora 4:370–457]
Largest genus of flowering plants endemic to North America. See also Keckiella , Nothochelone.

Native

P. heterophyllus Lindl.

Perennial 25–150 cm, woody-branched below
Leaves generally cauline, 20–95 mm, linear to oblanceolate, (sub)entire; base tapered
Inflorescence glabrous to short-hairy
Flower: calyx 4.2–8 mm, lobes lanceolate-acuminate to (ob)ovate; corolla 23–40 mm, magenta to blue, glabrous; anther sacs 2.2–3 mm, dehiscing across common tip to 1/2–2/3 their length, inner margins long-hairy; staminode glabrous
Ecology: Grassland, chaparral, forest openings
Elevation: 50–1600 m.
Bioregional distribution: Northwestern California, Cascade Range Foothills, n Sierra Nevada Foothills, Sacramento Valley, Central Western California, Southwestern California.

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bioregional map for PENSTEMON%20heterophyllus being generated
 


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