Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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RUBIACEAE

MADDER FAMILY

Lauramay T. Dempster

Annual, perennial herb, shrub, vine, tree
Leaves generally opposite, entire; stipules generally on stem, sometimes leaf-like (then leaves apparently whorled and stipules considered leaves), adjacent pairs sometimes fused
Inflorescence: cyme, panicle, cluster, or flower solitary, generally terminal and ± axillary
Flower generally bisexual; calyx generally ± 4-lobed, sometimes 0; corolla generally radial, 4-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1, ± fused if 2
Fruit: 2 or 4 nutlets or a berry, drupe, or capsule
Genera in family: ± 500 genera, 6000 species: worldwide, especially tropical; many cultivated (including Coffea , coffee; Cinchona , quinine; many ornamental)
Reference: [Dempster 1979 Fl CA 4(2):1–47]

GALIUM

BEDSTRAW, CLEAVERS

Annual, perennial herb, sometimes ± shrubby, often ± dioecious, glabrous or hairy, often scabrous
Stem when young 4-angled
Leaves in whorls of 4 or more, including leaf-like stipules
Inflorescence: panicle, or axillary clusters of 1–many flowers
Flower bisexual or unisexual (with sterile stamens or pistils); calyx 0; corolla generally rotate, sometimes ± bell-shaped, generally greenish, fading yellow or white, sometimes reddish, lobes generally 4; ovary 2-lobed, styles 2, ± fused basally
Fruit: 2 nutlets or 1 berry
Species in genus: ± 400 species: worldwide, especially temp
Etymology: (Greek: milk, from use of some species in its curdling)
Hairiness of ovary and fruit generally ± equal on a single plant; staminate plants often identified only by association with pistillate.

Native

G. trifidum L.

Perennial, rarely annual, minutely scabrous
Stem 10–50 cm, slender, weak, tangled
Leaves in whorls of 4–6, 4–19 mm, linear to elliptic or narrowly ovate-obovate, petioled; tip rounded
Inflorescence: few-flowered clusters; pedicel slender
Flower bisexual; corolla rotate, white or pinkish, generally 3-lobed
Fruit: nutlets, spheric, hard, smooth, black when dry; hairs 0
Ecology: Wet places
Elevation: < 2800 m.
Bioregional distribution: California Floristic Province, Great Basin Floristic Province
Distribution outside California: to Alaska, Rocky Mtns, Mexico

Native

var. pacificum Wiegand

Plant straggling
Stem 10–50 cm
Chromosomes: 2n=24
Ecology: Wet places
Elevation: < 2400 m.
Bioregional distribution: California Floristic Province, Great Basin Floristic Province
Distribution outside California: to Alaska, Mexico
Flowering time: Jul–Aug
Synonyms: var. subbiflorum Wiegand; G. cymosum Wiegand
Horticultural information: WET: 4, 5, 6, 17, 24 &SHD: 2, 3, 7, 8, 9, 14, 15, 16, 18, 19, 20, 21, 22, 23.

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bioregional map for GALIUM%20trifidum%20var.%20pacificum being generated
 
N.B. The distribution depicted here differs from that given in The Jepson Manual (1993)
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Galium trifidum var. pacificum
Retrieve dichotomous key for Galium
Overlay Consortium of California Herbaria specimen data by county on this map
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