Jepson Flora Project: Jepson Interchange    

 
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

RUBIACEAE

MADDER FAMILY

Lauramay T. Dempster

Annual, perennial herb, shrub, vine, tree
Leaves generally opposite, entire; stipules generally on stem, sometimes leaf-like (then leaves apparently whorled and stipules considered leaves), adjacent pairs sometimes fused
Inflorescence: cyme, panicle, cluster, or flower solitary, generally terminal and ± axillary
Flower generally bisexual; calyx generally ± 4-lobed, sometimes 0; corolla generally radial, 4-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1, ± fused if 2
Fruit: 2 or 4 nutlets or a berry, drupe, or capsule
Genera in family: ± 500 genera, 6000 species: worldwide, especially tropical; many cultivated (including Coffea , coffee; Cinchona , quinine; many ornamental)
Reference: [Dempster 1979 Fl CA 4(2):1–47]

GALIUM

BEDSTRAW, CLEAVERS

Annual, perennial herb, sometimes ± shrubby, often ± dioecious, glabrous or hairy, often scabrous
Stem when young 4-angled
Leaves in whorls of 4 or more, including leaf-like stipules
Inflorescence: panicle, or axillary clusters of 1–many flowers
Flower bisexual or unisexual (with sterile stamens or pistils); calyx 0; corolla generally rotate, sometimes ± bell-shaped, generally greenish, fading yellow or white, sometimes reddish, lobes generally 4; ovary 2-lobed, styles 2, ± fused basally
Fruit: 2 nutlets or 1 berry
Species in genus: ± 400 species: worldwide, especially temp
Etymology: (Greek: milk, from use of some species in its curdling)
Hairiness of ovary and fruit generally ± equal on a single plant; staminate plants often identified only by association with pistillate.

Native

G. oreganum Britton

OREGON BEDSTRAW

Perennial, erect, 20–30 cm, glabrous except leaf margin and veins
Leaves in whorls of 4, 20–40 mm, broadly ovate or elliptic, 3-nerved
Inflorescence: clusters, terminal on upper branches; flowers few
Flower bisexual: corolla rotate, yellowish or greenish
Fruit: nutlets; hairs long, hooked
Ecology: UNCOMMON. Open coniferous forest
Elevation: 1500 m.
Bioregional distribution: Klamath Ranges (Del Norte Co.)
Distribution outside California: w Oregon, Washington
See the CNPS Inventory for information about endangerment and rarity.
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bioregional map for GALIUM%20oreganum being generated
 


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