Jepson Flora Project: Jepson Interchange    

 
link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.

RUBIACEAE

MADDER FAMILY

Lauramay T. Dempster

Annual, perennial herb, shrub, vine, tree
Leaves generally opposite, entire; stipules generally on stem, sometimes leaf-like (then leaves apparently whorled and stipules considered leaves), adjacent pairs sometimes fused
Inflorescence: cyme, panicle, cluster, or flower solitary, generally terminal and ± axillary
Flower generally bisexual; calyx generally ± 4-lobed, sometimes 0; corolla generally radial, 4-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1, ± fused if 2
Fruit: 2 or 4 nutlets or a berry, drupe, or capsule
Genera in family: ± 500 genera, 6000 species: worldwide, especially tropical; many cultivated (including Coffea , coffee; Cinchona , quinine; many ornamental)
Reference: [Dempster 1979 Fl CA 4(2):1–47]

GALIUM

BEDSTRAW, CLEAVERS

Annual, perennial herb, sometimes ± shrubby, often ± dioecious, glabrous or hairy, often scabrous
Stem when young 4-angled
Leaves in whorls of 4 or more, including leaf-like stipules
Inflorescence: panicle, or axillary clusters of 1–many flowers
Flower bisexual or unisexual (with sterile stamens or pistils); calyx 0; corolla generally rotate, sometimes ± bell-shaped, generally greenish, fading yellow or white, sometimes reddish, lobes generally 4; ovary 2-lobed, styles 2, ± fused basally
Fruit: 2 nutlets or 1 berry
Species in genus: ± 400 species: worldwide, especially temp
Etymology: (Greek: milk, from use of some species in its curdling)
Hairiness of ovary and fruit generally ± equal on a single plant; staminate plants often identified only by association with pistillate.

Native

G. californicum Hook. & Arn.

CALIFORNIA BEDSTRAW

Perennial or sometimes ± woody above ground, low, tufted, or ± climbing, dioecious, ± hairy
Stem 5–90 cm
Leaves in whorls of 4, ± petioled, 3–20 mm, ovate to elliptic; tip acute to obtuse
Staminate inflorescence: clusters; flowers few
Pistillate inflorescence: flowers generally solitary in axils
Flower: corolla rotate, yellowish
Fruit: berry, ± hairy
Ecology: Moist, ± shaded sites, open slopes, forests, canyons, bluffs
Elevation: 15–1700 m.
Bioregional distribution: Outer North Coast Ranges, Inner North Coast Ranges, n Sierra Nevada Foothills, San Francisco Bay Area, Outer South Coast Ranges, n Channel Islands, Western Transverse Ranges, San Gabriel Mountains, San Jacinto Mountains.

Native

subsp. miguelense (Greene) Dempster & Stebbins

SAN MIGUEL ISLAND BEDSTRAW

Plant low, prostrate or climbing, dense, woody; hairs ± stout, curved
Stem 15–45 cm; internodes short
Leaf 2–8 mm, broadly ovate to round, leathery; margin ± turned down; tip acute to round, sometimes with small point
Ecology: UNCOMMON. Grassy slopes
Elevation: 30–60 m.
Bioregional distribution: n Channel Islands
Synonyms: var. m. (Greene) Jeps
See the CNPS Inventory for information about endangerment and rarity.
previous taxon | next taxon
bioregional map for GALIUM%20californicum%20subsp.%20miguelense being generated
 
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Galium californicum subsp. miguelense
Retrieve dichotomous key for Galium
Overlay Consortium of California Herbaria specimen data by county on this map
Show other taxa with the same California distribution | Read about bioregions | Get lists of plants in a bioregion
Return to the Jepson Interchange main page
Return to treatment index page
Glossary
    FEEDBACK
  • This page is no longer being maintained.


University & Jepson Herbaria Home Page |
General Information | University Herbarium | Jepson Herbarium |
Visiting the Herbaria | On-line Resources | Research |
Education | Related Sites
Copyright © by the Regents of the University of California