Jepson Flora Project: Jepson Interchange    

 
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RUBIACEAE

MADDER FAMILY

Lauramay T. Dempster

Annual, perennial herb, shrub, vine, tree
Leaves generally opposite, entire; stipules generally on stem, sometimes leaf-like (then leaves apparently whorled and stipules considered leaves), adjacent pairs sometimes fused
Inflorescence: cyme, panicle, cluster, or flower solitary, generally terminal and ± axillary
Flower generally bisexual; calyx generally ± 4-lobed, sometimes 0; corolla generally radial, 4-lobed; stamens epipetalous, alternate corolla lobes, generally included; ovary generally inferior, chambers generally 2 or 4, style 1, ± fused if 2
Fruit: 2 or 4 nutlets or a berry, drupe, or capsule
Genera in family: ± 500 genera, 6000 species: worldwide, especially tropical; many cultivated (including Coffea , coffee; Cinchona , quinine; many ornamental)
Reference: [Dempster 1979 Fl CA 4(2):1–47]

DIODIA

BUTTONWEED

Annual
Leaves opposite; stipules fused to leaf bases, small, fringed
Inflorescence: flowers 1–several in axils, sessile
Flower: corolla funnel-shaped, lobes generally 4
Fruit: 2 nutlets; calyx teeth persistent
Species in genus: 50 species: warm Am, Africa
Etymology: (Greek: thoroughfare, from habitats)

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