Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.


Barbara Ertter

Annual or perennial herb, sometimes monoecious; hairs 0 or partitioned, not glandular
Leaves alternate, odd-1-pinnate; leaflets generally evenly toothed or lobed
Inflorescence: spikes 1–many, head-like; peduncles long
Flowers bisexual (or upper pistillate, lower staminate); hypanthium urn-shaped; bractlets 0; sepals generally 4; petals 0; stamens 0–many; pistils 0–3, ovaries superior, continuous to style at top, stigma generally ± shrub-like, exserted
Fruit: hypanthium hard, 4-angled, enclosing achenes
Species in genus: ± 25 species: n temp, arctic
Etymology: (Latin: blood-absorbing, from styptic properties)
Reference: [Nordberg 1966 Opera Bot 11(2):1–103]


S. minor Scop. subsp. muricata Briq.


Perennial, tufted, taprooted
Stem erect, generally 2–7 cm
Leaves: basal present at flower, largest generally 4–20 cm; leaflets 4–10 per side, largest blade generally 5–20 mm, round-oblong, stalk generally 1–4 mm, teeth generally < 15, < 1/3 to midvein
Inflorescence 7–30 mm, 6–20 mm wide, ovoid-spheric, ± 5–30-flowered
Flower: sepals 3–6 mm, elliptic, green or purplish; stamens many, filaments thread-like
Fruit ± 5 mm; angles equally short-winged; faces with raised bumpy network
Chromosomes: n=14,28
Ecology: Open, especially disturbed areas
Elevation: 30–1600 m.
Bioregional distribution: California Floristic Province (except High Sierra Nevada)
Distribution outside California: to e US; native to Europe
Often used in seeding mixtures after fires and in pastures.

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bioregional map for SANGUISORBA%20minor%20subsp.%20muricata being generated

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