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ROSACEAE

ROSE FAMILY

Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.

HORKELIA

Barbara Ertter

Perennial, generally ± glandular, generally resinous-smelling; caudex generally branched
Stem generally ascending to erect
Leaves generally basal, odd-1-pinnate, generally ± flat; cauline alternate, reduced upward; uppermost lateral leaflets generally ± fused with terminal
Inflorescence: cyme, open or of dense clusters; pedicels generally straight
Flower: hypanthium a ± flat-bottomed cup, width ± 2 X length; bractlets 5, generally 2/3 sepals; sepals 5, often reflexed; petals 5, generally ± = sepals, blunt, white; stamens 10, filaments flat, often forming a tube; pistils 2–many, ovaries superior, styles jointed below fruit tip, ± thicker at base
Fruit: achene
Species in genus: 19 species: w North America
Etymology: (J. Horkel, German plant physiologist, 1769–1846)
Data apply to basal leaves, pressed hypanthia.

Native

H. wilderae Parish

BARTON FLATS HORKELIA

Plant ± rosetted, green; caudex generally simple
Stem generally decumbent, generally 10–25 cm
Leaf 4–10 cm; leaflets 4–7 per side, separated, 3–10 mm, ± obovate, divided 1/2–3/4 to base into 5–15 lobes, hairs ± sparse
Inflorescence open, generally 5–15-flowered; pedicels 3–15 mm, ± recurved in fruit
Flower: hypanthium width ± 2–3 mm, ± 2 X length; bractlets generally < 0.5 mm wide, lanceolate to ovate; sepals ± 2 mm; petals 2–3 mm, oblanceolate to oblong; filaments 0.5–1 mm, bases 0.5–1 mm wide, anthers ± 0.4 mm; pistils < 5, styles ± 1–1.5 mm
Fruit ± 2 mm
Ecology: Edge of chaparral, under pines
Elevation: 2200–3000 m.
Bioregional distribution: San Bernardino Mountains (Barton Flats area).

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