TREATMENT FROM THE JEPSON MANUAL	previous taxon | next taxon Jepson Interchange (more information)
	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ROSACEAE

ROSE FAMILY

Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.

HOLODISCUS

Richard A. Lis

Shrub, ± hairy
Stem 3–60 dm; bark reddish, in age gray, shredding
Leaves simple, alternate, 0.3–12 cm, thin to leathery, toothed; base truncate to generally ± wedge-shaped; lower surface strongly veined; stipules 0; petiole distinct or not
Inflorescence: panicle, ± terminal, dense, 2.5–25 cm, ± conic, many-flowered, persistent; pedicel slender, bractlets 1–3
Flower: hypanthium 3–5 mm wide, saucer-shaped, prominent nectary-disk below inner rim; sepals 5, 1–2 mm; petals 5, 1.5–2 mm, ± ovate, generally white; stamens 15–20, wider at base; pistils 5, ovaries superior, 2-ovuled, hairs dense, bristle-like, persistent in fruit, style 1 mm, persistent, stigma ± 2-lobed
Fruit: achenes 5, 1–1.5 mm, often with sessile glands
Species in genus: 5 species: w North America, C.Am, n South America
Etymology: (Greek: whole disk)
Spp. highly variable; leaves of peg-like stems best for identification; complexity in c SNH evidently from local climatic variation, hybridization.

Native

H. discolor (Pursh) Maxim.

OCEANSPRAY

Plant 1.5–6 m, ± open
Stem hairy; glands 0; peg-like stems not very predominant
Leaves of peg-like stems generally 2–4 cm, elliptic, others 2–12 cm, ovate to wide-ovate; teeth generally below middle of blade, generally of 2 sizes; petiole generally distinct, generally not winged
Inflorescence generally not mixed with leaves, 10–25 cm, 5–25 cm wide
Flower: sepal outer surface hairs generally sparse to dense, short, inner surface generally glabrous
Ecology: Moist woodland edges, rocky slopes
Elevation: < 1800 m.
Bioregional distribution: Northwestern California, High Sierra Nevada, Great Central Valley (Sutter Buttes), Central Western California, Channel Islands, Western Transverse Ranges, San Gabriel Mountains (300–1300 m), Peninsular Ranges
Distribution outside California: to British Columbia, Montana, Colorado, Texas, Mexico
Synonyms: var. delnortensis Ley; var. dumosus (S. Watson) Dippel; var. franciscanus (Rydb.) Jeps.; H. boursieri (Carrière) Rehder
Horticultural information: DRN: 4, 5, 6, 15, 16, 17 &SHD: 18, 24 &IRR: 1, 2, 3, 7, 8, 9, 14, 19, 20, 21, 22, 23; CVS.