Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.


Barbara Ertter

Annual, inconspicuous, nonglandular
Stem spreading to erect
Leaves palmately lobed
Inflorescence: flowers several per leaf axil, ± hidden by sheathing stipules
Flower: hypanthium ± urn-shaped; bractlets 0–4; sepals 4; petals 0; stamen generally 1; pistil generally 1, ovary superior, jointed to style near base
Fruit: hypanthium encasing achene
Species in genus: 10–20 species: worldwide, especially Medit
Etymology: (Greek: unseen, from hidden flowers)
Reference: [Rothmaler 1937 Feddes Repert 42:164–173]


A. occidentalis (Nutt.) Rydb.

Stem generally 2–10 cm
Leaf generally 3–12 mm; stipules widely ovate, deeply few-toothed; petiole (between stipule, blade) generally 1–5 mm; blade generally 2–5 mm, ± round, lobes 3, > 2/3 to base, again toothed or lobed
Flower 0.5–2 mm; bractlets 0–0.5 mm; sepals 0.2–0.6 mm
Fruit: achene ± 1 mm, ovoid
Ecology: Seasonally moist grassland, chaparral, woodland
Elevation: 30–1200 m.
Bioregional distribution: Northwestern California, Sierra Nevada Foothills, Sacramento Valley, Central Western California, Southwestern California
Distribution outside California: to Washington, Baja California
Synonyms: Alchemilla o. Nutt
Highly variable, with several ± separable forms. Relation to species of Medit unclear. n NCo plants with more hidden flowers resemble A. microcarpa (Boiss. & Reut.) Rothm. of eastern North America, Eur. A. arvensis L. (larger plants) weedy in OR, WA.

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bioregional map for APHANES%20occidentalis being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Aphanes occidentalis
Retrieve dichotomous key for Aphanes
Overlay Consortium of California Herbaria specimen data by county on this map
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