Jepson Flora Project: Jepson Interchange    

link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
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Annual to tree
Leaves simple to pinnately to palmately compound, generally alternate; stipules free to fused, persistent to deciduous
Inflorescence: cyme, raceme, panicle, or flowers solitary
Flower generally bisexual, radial; hypanthium free or fused to ovary, saucer- to funnel-shaped, often with bractlets alternate with sepals; sepals generally 5; petals generally 5, free; stamens (0)5–many, pistils (0)1–many, simple or compound; ovary superior to inferior, styles 1–5
Fruit: achene, follicle, drupe, pome, or blackberry- to raspberry-like
Seeds generally 1–5
Genera in family: 110 genera, ± 3000 species: worldwide, especially temp. Many cultivated for ornamental and fruit, especially Cotoneaster , Fragaria , Malus , Prunus , Pyracantha, Rosa , and Rubus
Reference: [Robertson 1974 J Arnold Arbor 55:303–332,344–401,611–662]
Family description, key to genera by Barbara Ertter and Dieter H. Wilken.


Barbara Ertter

Perennial, nonglandular
Leaves generally ± basal, odd-1-pinnate; leaflets ± evenly toothed to lobed
Inflorescence: dense spike or head
Flower: hypanthium ± conic; bractlets 0; sepals generally 4; petals 0; stamens 2 or 4, opposite sepals; pistils 1(2), ovaries superior, continuous to style at top, stigma many-branched, exserted from hypanthium
Fruit: hypanthium ± hardened, encasing elliptic achenes; prickles generally 4–many, barbed
Species in genus: 50–150 species: especially s hemisphere
Etymology: (Greek: thorn, from fruit)
Reference: [Yeo 1973 Bot Soc British Isles Conf Rep 13:193–221]


A. pinnatifida Ruiz & Pav. var. californica (Bitter) Jeps.

Stem: flower-stem 10–60 cm
Leaf 3–12 cm; stipules 0; leaflets 5–8 per side, 4–15 mm, ± oblanceolate-ovate, lobed nearly to midvein into 3–8 linear-lanceolate segments
Inflorescence: spike 10–20 mm diam, ± oblong
Flower: sepals 2.5–3.5 mm, elliptic-lanceolate; stamens generally 4, purple-black
Fruit: hypanthium body 4–7 mm, obovate, ± ridged; prickles > 4, scattered, generally 1–3 mm
Ecology: Coastal grassland, open, rocky slopes
Elevation: 50–400 m.
Bioregional distribution: s North Coast, Central Coast, w San Francisco Bay Area
Synonyms: A. c. Bitter
South America plants (var. p.) variable
Horticultural information: DRN: 4, 5, 15, 16, 17 &IRR: 14, 19, 20, 21, 22, 23, 24.

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bioregional map for ACAENA%20pinnatifida%20var.%20californica being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).

Retrieve Jepson Interchange Index to Plant Names entry for Acaena pinnatifida var. californica
Retrieve dichotomous key for Acaena
Overlay Consortium of California Herbaria specimen data by county on this map
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