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PORTULACACEAE

PURSLANE FAMILY

Annual or perennial herb, generally fleshy
Stems generally glabrous
Leaves simple, alternate or opposite, sometimes stipuled
Inflorescence various
Flower bisexual, radial; sepals generally 2(–8), free or fused at base; petals 3–18, free or ± fused; stamens 1–many, free or inserted on corolla; ovary superior or partly inferior, chamber 1, placenta free-central or basal; styles 2–8, generally fused at base
Fruit: capsule, circumscissile or 2–3-valved
Seeds 1–many, generally black, generally shiny
Genera in family: ± 20 genera, ± 400 species: generally temp Am, Australia, s Africa; some cultivated (Lewisia, Portulaca, Calandrinia )
Reference: [Bogle 1969 J Arnold Arbor 50:566–598]
Family description and key to genera by Dieter H. Wilken & Walter A. Kelley.

CLAYTONIA

Kenton L. Chambers

Annual or perennial herb, from stolon, rhizome, tuber, or taproot, glabrous, ± fleshy
Leaves entire; basal 0–many, rosetted; cauline generally 2, generally opposite, free to fully fused into ± 2-toothed disk or cup surrounding stem
Inflorescence: raceme, terminal, 1-sided; pedicels reflexed, becoming erect in fruit
Flower: petals 5, pink or white; stamens 5, epipetalous; ovary chamber 1, placentas basal, style 1, stigmas 3
Fruit: capsule; valves 3, margins rolling inward and forcibly expelling seeds
Seeds 3–6, generally black, generally clearly appendaged
Species in genus: 28 species: North America, e Asia
Etymology: (John Clayton, colonial Am botanist, born 1686)
Reference: [Miller 1978 Syst Bot 3:322–341; Miller & Chambers 1993 Novon 3:268–273]
Some species formerly placed in Montia.

Native

C. rubra (Howell) Tidestr.

Annual
Stem 1–15 cm
Leaves: basal 1–8 cm, blade < 2 cm, elliptic to widely deltate, base truncate to wedge-shaped, tip generally obtuse, petiole linear; cauline pair fused or partly free on 1 side, < 4 cm wide, generally round or with 2 squarish corners
Inflorescence sessile, ± dense, 1-bracted at base; flowers 3–30
Flower: sepals 1.5–3 mm; petals 2–3.5 mm, white or pinkish
Fruit 2–3 mm
Seed 1–2 mm, elliptic, shiny, smooth
Ecology: Vernally moist dunes, coniferous forest, scrub, in sun or shade
Elevation: < 2500 m.
Bioregional distribution: Northwestern California, High Cascade Range, High Sierra Nevada, Tehachapi Mountain Area, w Central Western California, Great Basin Floristic Province
Distribution outside California: to British Columbia, South Dakota, Colorado
Intergrades with C. parviflora , C. perfoliata.

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