|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL (1993)||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
AND IS MAINTAINED FOR ARCHIVAL PURPOSES ONLY
Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)45; stamens 45, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):128. See glossary p. 25 for illustrations of general family characteristics.
Annual to perennial herb (sometimes short-lived, dying after flowering once)
Stem generally erect
Leaves: lower generally tapered or petioled, often wavy-margined, generally pinnately lobed, ± dentate, lobes and teeth spine-tipped, margin generally spiny-ciliate, glabrous to tomentose; upper generally sessile, ± reduced
Inflorescence: heads discoid, 1many; involucre cylindric to spheric; phyllaries many, graduated in several series, outer spine-tipped; receptacle flat, long-bristly
Flowers generally many; corollas ± bilateral, white to red or purple, tube long, slender, lobes linear; anther bases sharply sagittate, tips oblong; style tip with slightly swollen node, appendage (above node) long, cylindric, branches very short
Fruit ovoid, glabrous; scar slightly angled; pappus bristles many, plumose, ± persistent or falling in ring
Species in genus: ± 200 species: North America, Eurasia
Etymology: (Greek: thistle)
Taxa difficult, incompletely differentiated, hybridize.
Biennial (or short-lived perennial herb) 110 dm, ± fleshy, bush-like or low, mounded, gray-tomentose throughout; hairs appressed, felt-likeSee the CNPS Inventory for information about endangerment and rarity.
Leaves generally very strongly wavy; lower 1025 cm, wing-petioled, entire or ± widely lobed, lobes entire or few-toothed, main spines 14 mm; cauline gradually reduced, well-distributed, upper sessile, clasping, ± lobed, often spinier than lower, spines < 8 mm
Inflorescence: heads 1several, ± clustered, closely subtended by uppermost leaves; peduncles 07 cm; involucres 34 cm, 46 cm diam, hemispheric or bell-shaped; phyllaries linear-lanceolate, persistently tomentose, outer and middle spiny-ciliate, spine tips 25 mm
Flowers: corollas 2025 mm, white to pale yellow, tube 1113 mm, throat 56 mm, lobes 58 mm; anthers brown
Fruit 57 mm, light-brown to black, thick or ± compressed; pappus 1520 mm
Ecology: Dunes, bluffs
Elevation: < 20 m.
Bioregional distribution: s Central Coast (s San Luis Obispo and n Santa Barbara cos.).