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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.



David J. Keil and Charles E. Turner

Annual to perennial herb (sometimes short-lived, dying after flowering once)
Stem generally erect
Leaves: lower generally tapered or petioled, often wavy-margined, generally pinnately lobed, ± dentate, lobes and teeth spine-tipped, margin generally spiny-ciliate, glabrous to tomentose; upper generally sessile, ± reduced
Inflorescence: heads discoid, 1–many; involucre cylindric to spheric; phyllaries many, graduated in several series, outer spine-tipped; receptacle flat, long-bristly
Flowers generally many; corollas ± bilateral, white to red or purple, tube long, slender, lobes linear; anther bases sharply sagittate, tips oblong; style tip with slightly swollen node, appendage (above node) long, cylindric, branches very short
Fruit ovoid, glabrous; scar slightly angled; pappus bristles many, plumose, ± persistent or falling in ring
Species in genus: ± 200 species: North America, Eurasia
Etymology: (Greek: thistle)
Taxa difficult, incompletely differentiated, hybridize.


C. ciliolatum (L.F. Hend.) J.T. Howell


Perennial 6–20 dm; rootstock branched
Stem generally simple below, ± cobwebby to densely white-tomentose
Leaves gray-tomentose above, ± white below: rosettes from rootstock branches; lowest 1–2.5 dm, wing-petioled, entire to shallowly lobed, spines 0 or 1–2 mm; cauline well developed, gradually reduced, short-decurrent as spiny wings, shallowly to deeply lobed, weakly spiny, uppermost bract-like
Inflorescence: heads few–many in ± raceme- or panicle-like clusters; peduncles generally 1–10 cm; involucres ± 2 cm, 2–5 cm diam, ovoid to hemispheric, thinly tomentose; phyllaries ± linear, tips ascending to spreading, midribs of outer and middle with sticky glandular area, spine tips 1–3 mm, inner with tips flat, straight
Flowers: corollas 17–25 mm, dull white to lavender, tube 7–11 mm, throat 5–7 mm, lobes 5–7 mm
Fruit 3.5–4.5 mm, straw-colored to brown, thick; pappus ± 15 mm
Ecology: Open woodland
Elevation: 1000–1400 m.
Bioregional distribution: Klamath Ranges
Distribution outside California: s Oregon
See the CNPS Inventory for information about endangerment and rarity.
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