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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

CHRYSOTHAMNUS

RABBITBRUSH

Loran C. Anderson

Perennial to shrub
Stems erect, often densely clustered
Leaves alternate, sessile, entire
Inflorescence: heads discoid, arrayed in ± dense cymes, peduncled or subsessile; involucre generally cylindric; phyllaries generally in 3–5 series (± 5 vertical ranks), free, overlapping, keeled; receptacle naked
Flowers 2–20 (often 5) per head; corollas generally yellow, lobes 0.5–3 mm, generally spreading; style branches long, slender, generally exserted
Fruit narrowly cylindric, ± 5-ridged, generally light brown; pappus of many white to brownish bristles
Species in genus: 16 species: sw Can to n Mex
Etymology: (Greek: golden shrub)
Closely related to (perhaps part of) Ericameria.

Native

C. parryi (A. Gray) Greene

Subshrub or shrub, 1–9 dm
Stems prostrate to erect, white to green, tomentose
Leaves 1–7 cm, generally < 4 mm wide, thread-like to ± oblanceolate, ± glabrous to tomentose
Inflorescence: heads generally ± many, in long or rounded cymes; involucre 10–18 mm, widely cylindric; phyllaries ± lanceolate, weakly 5-ranked, keeled, ± membranous, tips green
Flowers 5–18; corollas 8–12.5 mm; style appendage > stigma
Fruit 4–7 mm, hairy; pappus generally < corolla
Chromosomes: 2n=18
Ecology: Dry places, generally open forest
Elevation: 700–3700 m.
Bioregional distribution: Klamath Ranges, North Coast Ranges, Cascade Range, High Sierra Nevada, San Bernardino Mountains, Great Basin Floristic Province, Desert Mountains
Distribution outside California: to s Oregon, Colorado, Arizona
12 subspp. in w US.

Native

subsp. imulus H.M. Hall & Clem.

Leaves 1–2 cm, spreading to recurved, green; glands sessile
Inflorescence: cyme, dense; phyllaries strongly recurved, yellowish green, ± resinous
Flowers 10–18
Ecology: Uncommon. Dry flats, open yellow-pine forest
Elevation: ± 2200 m.
Bioregional distribution: San Bernardino Mountains (Bear Valley). Perhaps only an extreme form of subsp. asper.