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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

BLEPHARIZONIA

BIG TARWEED


Species in genus: 1 sp
Etymology: (Greek: eyelash girdle, from fruits weakly held by phyllaries)

Native

B. plumosa (Kellogg) Greene

Annual 3–18 dm, strongly scented
Stem generally 1, erect, much-branched, soft-hairy to bristly, puberulent, ± glandular
Leaves basal and lower cauline 4–15 cm, crowded; blades linear, entire or serrate, often withering in flower; cauline alternate, generally subtending smaller leaf clusters, base 1–3 cm wide; clustered leaves < cauline, margins with tack-shaped glands
Inflorescence ± glandular; heads radiate, axillary or terminal, in a narrow to open panicle; bracts generally glandular-ciliate; involucre 5–7.5 mm, hemispheric; phyllaries 1 per ray flower, elliptic, hairy and often glandular, base closely appressed to ovary, tip free, acute; chaff scales between ray and disk flowers, = phyllaries, flat
Ray flowers 5–13; corolla 6–9 mm; ligule 5–8 mm, white, red-veined
Disk flowers 10–35; corolla 4–5.5 mm, white, throat > tube, lobes 0.5 mm; anthers brown; style branches tapered, short-bristly
Fruit 3.5–4 mm, club-shaped, 10-ribbed, densely puberulent, ± dark; ray pappus 0 or minute crown, sometimes of narrow, plumose scales < 1 mm; disk pappus 1.5–3 mm, scales 15–20, narrow, plumose, sometimes 0 or reduced to crown
Chromosomes: 2n=28
Ecology: Dry grassy areas
Elevation: < 1100 m.
Bioregional distribution: w San Joaquin Valley, e San Francisco Bay Area, South Coast Ranges.

Native

subsp. plumosa

Plant gray-green, sparsely glandular below heads
Inflorescence: branches of older plants spike-like or raceme-like; many heads lateral on inflorescence branches; involucre canescent
Fruit: disk pappus 1.5–3 mm
Ecology: Dry hills, plains
Elevation: < 400 m.
Bioregional distribution: nw San Joaquin Valley (n of Alameda Co.), e San Francisco Bay Area.

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