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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

ARNICA

Theodore M. Barkley

Perennial generally from long, naked rhizome
Leaves: basal 0 or generally withered by flower; cauline opposite
Inflorescence ± flat-topped; heads radiate or discoid, 1–many; involucre hemispheric to obconic; phyllaries generally in 2 ± equal series; receptacle ± flat, naked
Ray flowers (0)6–21; ligules (orange-)yellow
Disk flowers many; corolla generally soft-hairy, colored like ligules; anther bases entire or slightly sagittate, tips triangular; style branches flat, tips truncate, very short, hair-tufted
Fruit ± cylindric, 5–10-veined; pappus of many barbed to subplumose bristles, white to red-brown
Species in genus: ± 27 species: North America, Eurasia
Etymology: (Latin or Greek: ancient name)
Reference: [Downie & Denford 1988 Rhodora 90:245–275]
Diploid species sexual; polyploid species generally form seeds asexually.

Native

A. chamissonis Less. subsp. foliosa (Nutt.) Maguire

Plant 3–8 dm, ± hairy, generally glandular upward
Stem 1, branched above
Leaves: cauline 5–10 pairs, sessile or short-petioled, 5–20(30) cm, blade (ob)lanceolate, (sub)entire, upper ± reduced
Inflorescence: heads radiate, 5–10; involucre 8–15 mm, bell-shaped; phyllaries ± obtuse, inner face generally hair-tufted near tip
Ray flowers generally 13; ligules 1–2 cm
Fruit 4–5 mm, ± glabrous to short-hairy, glandular; pappus short-barbed or weakly subplumose, dirty white to straw-colored
Chromosomes: 2n=38,57,106–108
Ecology: Damp meadows, rocky places, coniferous forest
Elevation: 1800–3500 m.
Bioregional distribution: Klamath Ranges, Cascade Range, Sierra Nevada, San Bernardino Mountains, Warner Mountains, n East of Sierra Nevada, White and Inyo Mountains
Distribution outside California: to British Columbia, Montana, New Mexico
Flowering time: Jul–Aug
Synonyms: vars. bernardina (Greene) Maguire, incana (A. Gray) Hultén, jepsoniana Maguire
Generally asexual
Horticultural information: IRR, SHD: 1, 2, 3, 4, 5, 6, 7, 10, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24.