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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

ADENOPHYLLUM

Annual, perennial herb, subshrubs
Stems erect
Leaves simple or pinnate, opposite or alternate, dotted with embedded oil glands
Inflorescence: heads radiate or discoid, solitary or in few-headed, often leafy-bracted, often open cymes; peduncles slender, with leaf-like or scale-like bracts; involucre bell-shaped to cylindric; phyllaries in 3 series, outer << others, free, 2 inner ± equal, fused or free, gland-dotted; receptacle flat, bearing short, fringed scales
Ray flowers 0–few; corollas yellow to red
Disk flowers few–many; corollas yellow to orange; style tips tapered
Fruit obconic, ribbed; pappus of scales dissected into slender bristles
Species in genus: 10 species: sw US, n Mex
Etymology: (Greek: gland-leaf)
Reference: [Strother 1986 Sida 11:371–378]

Native

A. porophylloides (A. Gray) Strother

Subshrub, glabrous; odor unpleasant
Stems many, 2–6 dm, generally much-branched
Leaves opposite below, alternate above, 1.5–4 cm, deeply divided; lobes linear, entire to sharply serrate, each with 1 gland at base, 1 near tip
Inflorescence: heads radiate (discoid); peduncles 2–8 cm, naked or with 1–5 narrow bracts; involucre cylindric; outer phyllaries 12–16, 3–8 mm, erect or recurved, each with a central gland; inner phyllaries 12–20, 10–15 mm, lanceolate, fused below, gland-dotted
Ray flowers (0)8–12; ligules 2–4 mm, yellow to red-orange
Disk flowers many; corollas 7–8 mm, yellow-orange
Fruit 5 mm, sparsely hairy; pappus scales 8–12, 7–8 mm, each dissected into 7–11 bristles
Chromosomes: 2n=26
Ecology: Dry, rocky hillsides, washes
Elevation: 200–1460 m.
Bioregional distribution: s Mojave Desert, Sonoran Desert
Distribution outside California: to Arizona, n Mexico
Flowering time: Mar–Jun
Synonyms: Dyssodia p. A. Gray