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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

TONESTUS

Gregory K. Brown

Perennial from taproot or branched caudex, glandular
Leaves: basal persistent; cauline alternate, reduced above, oblanceolate, 1–3-veined
Inflorescence loose; heads radiate, 1–5; involucre bell-shaped; phyllaries in 3–4 ± equal series, lanceolate, 1-veined, outer graduated into upper leaves; receptacle convex, naked
Ray flowers 10–25; ligules < 12 mm, yellow
Disk flowers many; corolla flaring slightly, yellow
Fruit narrowly oblong, ± cylindric, fusiform, or compressed, hairy; pappus of many white bristles in 1 series
Species in genus: ± 4 species: w North America
Etymology: (Anagram of Stenotus)
Reference: [Nesom & Morgan 1990 Phytologia 68:174–180]

Native

T. lyallii (A. Gray) A. Nelson

LYALL'S HAPLOPAPPUS


Stems 5–15 cm
Leaves < 6 cm, entire
Inflorescence: involucre 8–11 mm; phyllaries green to reddish
Disk flowers: corollas 6.5–7.5 mm
Fruit 3–4 mm; pappus = disk corolla
Chromosomes: 2n=18
Ecology: Alpine talus, barrens
Elevation: 2500–2700 m.
Bioregional distribution: c Klamath Ranges (Trinity Alps)
Distribution outside California: to British Columbia, Montana, Wyoming, Colorado
Synonyms: Haplopappus l. A. Gray

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