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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

RAILLARDIOPSIS

Bruce G. Baldwin

Perennial from caudex or woody rhizome, hairy to glandular-hairy
Stems ascending or erect, generally < or = 6 dm, often branched
Leaves opposite below, alternate above, simple, linear to lanceolate, entire, sessile
Inflorescence glandular; heads radiate or discoid, solitary or in leafy cymes, cylindric to bell-shaped; peduncles 0–8 cm; phyllaries 0–3, folded around ray ovaries; chaff scales in 1 involucre-like series, ± equal, ± fused
Ray flowers 0–3; corollas yellow; ligules deeply lobed; style branches long, ± glabrous
Disk flowers 5–29; corollas yellow; style branches long, bristly to base
Fruit ± cylindric, black; ray achenes curved, glabrous to partly hairy; disk achenes ± straight, ascending-hairy; pappus of flat, ciliate-plumose bristles
Species in genus: 2 species: montane CA-FP
Etymology: (Greek: like Raillardia or Railliardia )
Reference: [Baldwin & Kyhos 1990 Madroño 37:43–54]
Close to Madia and Hawaiian silversword alliance (including Railliardia ).

Native

R. muirii (A. Gray) Rydb.

MUIR'S RAILLARDELLA

Plant often matted; rhizome loosely branched
Leaves < or = 42 mm, greenish, not clasping; tip acute
Inflorescence: chaff scales 5–16, 8–13 mm, glandular-hairy on outer surface
Ray flowers 0
Disk flowers 7–29; corollas 6.5–10 mm
Fruit 3.5–7.5 mm, 0.5–1.2 mm wide, linear; pappus bristles 9–17, = or > mature achene
Chromosomes: 2n=16
Ecology: Dry, open sites on granitic soil
Elevation: 1100–2500 m.
Bioregional distribution: s High Sierra Nevada, Outer South Coast Ranges (Ventana Double Cone, Monterey Co.).
Synonyms: Raillardella m. A. Gray
Recent taxonomic note: *Carlquistia muirii (A. Gray) B.G. Baldwin

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