This page is based on the 1993 Jepson Manual.
Please see the Jepson eFlora for up-to-date information about California vascular plants.
|Jepson Flora Project: Jepson Interchange|
|TREATMENT FROM THE JEPSON MANUAL||
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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
Print edition is available from the University of California Press
|The second edition of The Jepson Manual (2012) is available from the University of California Press|
|See also the Jepson eFlora, which parallels the Second Edition|
Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)45; stamens 45, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):128. See glossary p. 25 for illustrations of general family characteristics.
Annual or perennial herb, generally densely glandular, aromatic
Stems 1several, generally simple below, ± branched above
Leaves generally opposite below, alternate above, generally linear to lanceolate, entire to slightly toothed
Inflorescence: heads generally radiate, generally peduncled, fewmany; phyllaries generally 120, free, enclosing (and falling with) ray achenes; receptacle ± flat, generally glabrous; chaff scales generally ± fused, in ring between ray and disk flowers
Ray flowers generally 120, sometimes minute; ligules 23-lobed, generally yellow
Disk flowers 1many, sometimes staminate; corollas yellow or maroon; anther tips triangular-ovate; style tips linear to oblong, acute, bristly
Fruit club-shaped or obovoid; ray achenes compressed, thickened, or 3-angled (1 angle toward center of head), ridged, sometimes beaked; pappus 0 or of short scales; disk achenes ± symmetric; pappus 0 or of 410 scales or bristles
Species in genus: 21 species: w North America, sw South America
Recent taxonomic note: *See revised taxonomy of Baldwin 1999 Novon 9:462471.
Etymology: (Chilean name)
Reference: [Nelson & Nelson 1980 Brittonia 32:323325]
Annual 125 dm, strongly scented
Stems simple to branched throughout, often very leafy below, soft-hairy below, sparsely to densely stalked-glandular especially above; glands yellow to black
Leaves 220 cm, linear to widely lanceolate, entire or ± serrate, soft-hairy to bristly, ± glandular
Inflorescence: heads in open, rounded to ± flat-topped cymes (sometimes panicle-like); involucre 4.512 mm, bell-shaped to hemispheric; phyllaries ± bristly, glandular or not, tips flat; receptacle hairy; chaff scales strongly fused
Ray flowers 521; corolla tubes 0.51 mm, ligules (2.5)420 mm, fan-shaped, deeply lobed, yellow, base generally maroon-spotted
Disk flowers 2550+, staminate; corollas 2.55 mm, yellow or maroon; anthers yellow or black
Fruit: ray achenes 2.55 mm, compressed side-to-side or ± 3-angled, glabrous, black or dark brown, sometimes mottled, beak 0, pappus 0; disk ovaries slender, much reduced, pappus 0
Ecology: Grassland, open forest
Elevation: < 3350 m.
Bioregional distribution: California Floristic Province (except Channel Islands), Great Basin Floristic Province
Distribution outside California: to n Oregon, Baja California
Highly variable; intermediates blur distinctions among extremes.
|YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).|