Jepson Flora Project: Jepson Interchange    

link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.



Annual or perennial herb, generally densely glandular, aromatic
Stems 1–several, generally simple below, ± branched above
Leaves generally opposite below, alternate above, generally linear to lanceolate, entire to slightly toothed
Inflorescence: heads generally radiate, generally peduncled, few–many; phyllaries generally 1–20, free, enclosing (and falling with) ray achenes; receptacle ± flat, generally glabrous; chaff scales generally ± fused, in ring between ray and disk flowers
Ray flowers generally 1–20, sometimes minute; ligules 2–3-lobed, generally yellow
Disk flowers 1–many, sometimes staminate; corollas yellow or maroon; anther tips triangular-ovate; style tips linear to oblong, acute, bristly
Fruit club-shaped or obovoid; ray achenes compressed, thickened, or 3-angled (1 angle toward center of head), ridged, sometimes beaked; pappus 0 or of short scales; disk achenes ± symmetric; pappus 0 or of 4–10 scales or bristles
Species in genus: 21 species: w North America, sw South America
Recent taxonomic note: *See revised taxonomy of Baldwin 1999 Novon 9:462–471.
Etymology: (Chilean name)
Reference: [Nelson & Nelson 1980 Brittonia 32:323–325]


M. bolanderi (A. Gray) A. Gray

Perennial 5–12 dm, strongly scented
Stems ± bristly below; upper branches densely stalked-glandular; glands dark brown to black
Leaves 10–30 cm, opposite well up stem, fused at base, bristly; upper alternate, often much reduced
Inflorescence: heads few, long-peduncled, in rounded or flat-topped clusters; involucre 9–13 mm, bell-shaped or hemispheric; phyllaries densely glandular, ± bristly, tips acuminate; chaff scales free or fused only near base
Ray flowers 8–12, showy; corolla tubes ± 3 mm, ligules 7–13 mm
Disk flowers 30–65, fertile; corollas 5–7 mm; anthers black
Fruit: ray achenes 5–7 mm, club-shaped, moderately curved, ± compressed side-to-side, obscurely 5-veined, dark brown or black, ± glabrous, beak ± 0, pappus 0 or minute crown; disk achenes 5–8 mm, narrowly club-shaped, short-hairy, brown; pappus scales 5–10, 1–5 mm, linear-lanceolate, ciliate
Chromosomes: 2n=12
Ecology: Meadows, streambanks
Elevation: 1000–2600 m.
Bioregional distribution: Klamath Ranges, High North Coast Ranges, High Cascade Range, High Sierra Nevada
Distribution outside California: s Oregon
Recent taxonomic note: *Kyhosia bolanderi (A. Gray) B.G. Baldwin

previous taxon | next taxon
bioregional map for MADIA%20bolanderi being generated

Retrieve Jepson Interchange Index to Plant Names entry for Madia bolanderi
Retrieve dichotomous key for Madia
Retrieve multiple-entry key (MEKA) for Madia
Overlay Consortium of California Herbaria specimen data by county on this map
Show other taxa with the same California distribution | Read about bioregions | Get lists of plants in a bioregion
Return to the Jepson Interchange main page
Return to treatment index page

University & Jepson Herbaria Home Page |
General Information | University Herbarium | Jepson Herbarium |
Visiting the Herbaria | On-line Resources | Research |
Education | Related Sites
Copyright © by the Regents of the University of California