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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

HULSEA

Dieter H. Wilken

Annual to perennial herb
Stems 1–5, 1–15 dm, ± hairy, glandular
Leaves alternate; petioles generally ciliate; blades generally ± oblanceolate, entire to lobed, ± reduced upward
Inflorescence raceme-like; heads radiate, 1–many; bracts ± narrowly lanceolate; involucre hemispheric to obconic; phyllaries many, in 2–3 series, linear to obovate, green, ± glandular; receptacle naked
Ray flowers: ligules yellow to red
Disk flowers many; corollas 5–9 mm, yellow to orange, generally glabrous
Fruit 4–10 mm, linear to club-like, black, ± hairy; pappus scales generally 2 pairs, 4–10 mm, generally deeply cut, generally translucent
Species in genus: ± 8 species: w US
Etymology: (G.W. Hulse, US Army surgeon, botanist, 1807–1883)
Reference: [Wilken 1977 Madroño 24:48–55]
Self-sterile.

Native

H. vestita A. Gray

Perennial generally 1–10 dm
Stems generally leafy in lower 1/3–1/2
Leaves: basal < 8 cm, 1–3 cm wide, spoon-shaped, entire to lobed, woolly above; cauline few
Inflorescence: heads < 15 mm, < 12 mm wide; bracts lanceolate to ovate, glandular, ± long-soft-hairy; phyllaries 8–11 mm, oblong to obovate, acuminate, hairy
Ray flowers 9–32; corollas 5–18 mm, 2–5 mm wide, yellow to red, puberulent
Fruit 5–7 mm, moderately hairy; pappus 1–2 mm, generally ± equal
Ecology: Open gravel, talus slopes
Elevation: 1300–3900 m.
Bioregional distribution: c&s High Sierra Nevada, Transverse Ranges, Peninsular Ranges, East of Sierra Nevada, Desert Mountains
Distribution outside California: Nevada
Sspp. generally geog separated.

Native

subsp. vestita

Plant 1–4 dm
Stems sometimes ± leafless
Leaves: petiole = blade, green; basal leaves entire, woolly
Inflorescence: bracts densely woolly; phyllary tips green or red-tinged
Flowers generally yellow; ray corollas 5–9 mm
Chromosomes: 2n=38
Ecology: Montane sagebrush to fir forest
Elevation: 2400–3000 m.
Bioregional distribution: c&s High Sierra Nevada, n East of Sierra Nevada
Flowering time: May–Jul

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