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Jepson Interchange (more information)
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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.


Bruce G. Baldwin and Susan J. Bainbridge

Species in genus: 1 sp
Etymology: (Greek: whole girdle)
Reference: [Thompson 1983 PhD thesis Univ CA Davis]


H. filipes (Hook. & Arn.) Greene

Perennial 3–15 dm, from rhizomes, slender-branched, ± white-hairy and glandular above
Leaves < 10 cm, linear to lanceolate; glands tack-shaped (especially margins); lower leaves opposite, fused around stem; upper leaves alternate, sessile
Inflorescence: heads radiate in leafy cyme; involucre obconic; peduncles < 6 cm; phyllaries 3–5 mm, each enfolding a ray fruit, long-white-hairy, often glandular; receptacle convex; chaff scales 8–13, between ray and disk flowers, hairy, edges fused
Ray flowers 6–10, fruiting; ligules 2.5–5 mm, deeply linear-lobed, white, purplish below, sparsely hairy; style branches long, glabrous
Disk flowers 9–28, staminate; corollas 3–4.5 mm, white, hairy; anthers purple, acute; style branches long, short-bristly
Fruit 3–5 mm, narrowly club-shaped, compressed front-to-back, glabrous; inner surface ribbed; ray pappus 0; disk pappus bristles 0–5
Chromosomes: 2n=28
Ecology: Dry beds and borders of lakes, streams
Elevation: < 1000 m.
Bioregional distribution: North Coast Ranges, Cascade Range Foothills, n&c Sierra Nevada Foothills, Great Central Valley (edge), San Francisco Bay Area, c Outer South Coast Ranges.

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