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ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

HESPEREVAX

James D. Morefield

Annual < 18 cm, ± green to gray-woolly
Leaves alternate or seeming whorled; base hard, yellowish; blade oblanceolate to ± round, entire
Inflorescence: heads disciform, sessile, 1–many above (sometimes scattered below), leafy-bracted, < 5 mm; phyllaries 0; receptacle conic, ± acuminate, short-shiny-bristly; chaff scales phyllary-like, subtending flowers, flat to concave, persistent, rigid, outer acute, ± glabrous, inner obtuse, finely woolly inside
Pistillate flowers in 1–5 series, cylindric
Disk flowers staminate, 2–6(12), ± bilateral, 3–5-lobed
Fruit obovoid, compressed front-to-back, ± angled, smooth, dull, ± black-banded near base; pappus 0
Species in genus: 3 species: CA, s OR
Etymology: (Greek: western Evax )
Reference: [Morefield 1992 Syst Bot 17:293–310]
Segregated from Evax ; closest to Ancistrocarphus. Evax multicaulis DC. and E. prolifera of AZ & NM not adequately documented from CA.

Native

H. caulescens (Benth.) A. Gray

HOGWALLOW STARFISH


Stems 0–few, erect, < 18 cm
Leaves ± crowded near stem tips, long-petioled, longest generally 33–90 mm, 7–20 mm wide
Inflorescence: upper heads in dense groups of 10–40 subtended by generally 10–20 leaves, 3–5 mm, 2.5–4 mm wide, largest group > 9 mm wide; lower heads 0; receptacle < 2 X longer than wide; chaff in 5 vertical ranks, inner scales 3–4 mm, tip exserted, spreading over outer scales
Pistillate flowers in 2–4 series
Disk flowers 1.1–1.6 mm
Fruit generally 1.5–2 mm
Ecology: Dry mud of vernal pools and flats
Elevation: 0–500 m.
Bioregional distribution: s Sierra Nevada Foothills, n&c Great Central Valley, sw Peninsular Ranges
Synonyms: Evax c. (Benth.) A. Gray; E. acaulis Greene (1894); E. a. (Kellogg) Greene (1883) misapplied
Tall, narrow-lvd plants have been called E. involucrata Greene. Becoming uncommon.

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