Jepson Flora Project: Jepson Interchange    

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Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California

  • Up-to-date information about California vascular plants is available from the Jepson eFlora.



David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.



Meredith A. Lane

Biennial to subshrub from taproot or woody caudex, glabrous to tomentose or glandular-sticky
Leaves entire to pinnately lobed, generally clasping, gland-dotted
Inflorescence: heads generally radiate, 1–many; involucres obconic to hemispheric, generally gummy; phyllaries in 4–10 series, bases generally tough, tips green; receptacle flat to convex, naked, ± pitted
Ray flowers 0–many; ligules yellow
Disk flowers: corollas yellow; style appendages linear to lanceolate, generally = or > stigmatic portion
Fruit cylindric or swollen-obconic, shiny white to ± brown, glabrous, smooth to ridged; pappus of 1–6 awns ± < disk corollas, generally < 0.2 mm wide, generally U-shaped in X -section, generally entire, deciduous
Species in genus: ± 80 species: c&w North America, South America
Etymology: (D.H. Grindel, 1776–1836, Latvian botanist)
Hybrids common.


G. hirsutula Hook. & Arn.

Perennial 2–15 dm, erect, few-branched above, green to red-purple or -brown, glabrous to tomentose
Leaf 1–10 cm, oblong to lanceolate, entire to lobed (less so upward) yellow-, red-, or gray-green
Inflorescence: heads often subtended by phyllary-like bracts; involucres 7–32 mm diam, hemispheric to bell-shaped; phyllaries 4–5 series, generally lanceolate-acute, outer erect to reflexed
Ray flowers 10–60+; ligules 8–20 mm
Disk flowers many; throat narrow
Fruit 2.5–5.5 mm, golden- to red-brown, smooth to ridged; pappus awns flat
Chromosomes: 2n=12,24
Ecology: Sandy, clay, or serpentine slopes or roadsides
Elevation: < 1700 m.
Bioregional distribution: North Coast Ranges, n&c Sierra Nevada Foothills, Sacramento Valley, Central Western California, Western Transverse Ranges, Peninsular Ranges, Sonoran Desert
Distribution outside California: to British Columbia?
Varieties (except hallii ) intergrade.

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