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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

ERIOPHYLLUM

WOOLLY SUNFLOWER

John S. Mooring and Dale E. Johnson

Annual to shrubs, ± woolly
Leaves generally alternate, entire to nearly compound
Inflorescence: heads 1–many, generally radiate; cluster often ± flat-topped; involucre obconic to hemispheric; phyllaries in 1 series, free or ± fused; receptacle flat to columnar, generally naked
Ray flowers generally ± 1 per phyllary; ligules entire to lobed, generally yellow (white)
Disk flowers (3)10–300; corolla yellow; anther tips ovate, deltate or awl-shaped
Fruit 4-angled or flattened in outer flowers, generally club-shaped in inner flowers; pappus 0–15 ± jagged or fringed scales
Species in genus: 14 species: w North America
Etymology: (Greek: woolly leaf)
Reference: [Mooring 1991 Madroño 38:213–226]
Annual species by Dale E. Johnson.

Native

E. pringlei A. Gray

Annual 1–8 cm, ± tufted-spreading, white-woolly
Leaf 3–10 mm, wedge-shaped, generally 3-lobed, very woolly; margins rolled under
Inflorescence: heads discoid, ± sessile in leafy clusters at branch tips; involucre 3–6 mm, hemispheric; phyllaries 6–8, acuminate, free; receptacle convex
Ray flowers 0
Disk flowers 10–25; corollas ± 2 mm, minutely glandular; anther tips deltate
Fruit 1.5–2 mm, strigose; pappus ± 1 mm
Chromosomes: 2n=14+
Ecology: Chaparral, sagebrush or desert scrub or woodlands
Elevation: 300–2200 m.
Bioregional distribution: s Sierra Nevada Foothills, Tehachapi Mountain Area, s Outer South Coast Ranges, Inner South Coast Ranges, Transverse Ranges, East of Sierra Nevada, Desert
Distribution outside California: to s Nevada, Arizona
Flowering time: Apr–Jun
Horticultural information: TRY.