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	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ASTERACEAE

SUNFLOWER FAMILY

David J. Keil, Family Editor and author, except as specified

Annual to tree
Leaves basal or cauline, alternate to whorled, simple to compound
Inflorescence: 1° inflorescence a head, each resembling a flower, 1–many, generally arrayed in cymes, generally subtended by ± calyx-like involucre; flowers 1–many per head
Flowers bisexual, unisexual, or sterile, ± small, of several types; calyx 0 or modified into pappus of bristles, scales, or awns, which is generally persistent in fruit; corolla radial or bilateral (rarely 0), lobes generally (0)4–5; stamens 4–5, anthers generally fused into cylinder around style, often appendaged at tips, bases, or both, filaments generally free, generally attached to corolla near throat; pistil 1, ovary inferior, 1-chambered, 1-seeded, style 1, branches 2, generally hair-tufted at tip, stigmas 2, generally on inside of style branches
Fruit: achene, cylindric to ovoid, generally deciduous with pappus attached
Genera in family: ± 1300 genera, 21,000 species (largest family of dicots): worldwide. Largest family in CA. Also see tribal key to CA genera: Strother 1997 Madroño 44(1):1–28. See glossary p. 25 for illustrations of general family characteristics.

ERICAMERIA

GOLDENBUSH

Gregory K. Brown and David J. Keil

Shrubs < 50 dm, resinous, generally gland-dotted
Leaves < 10 cm, thread-like to wedge-shaped, entire
Inflorescence various; heads radiate or discoid; involucre 3–14 mm, obconic to hemispheric; phyllaries in 2–6 series, ± lanceolate to ovate, generally resinous, tips erect to recurved, obtuse to acuminate or tailed, midrib often thickened with a resin gland
Ray flowers 0–30; corollas 2–12 mm, generally yellow
Disk flowers 4–70+; corollas 3–11 mm, yellow
Fruit 2–8 mm, ribbed; pappus white to brown
Species in genus: ± 27 species: w North America
Reference: [Nesom 1990 Phytologia 68:144–155]
Generally flowers summer/autumn. Some species hybridize with Chrysothamnus nauseosus.

Native

E. nana Nutt.

Plant 1–5 dm, glabrous
Leaf 10–15 mm, linear to narrowly oblanceolate, generally curved, acute, not gland-dotted, ± sticky
Inflorescence: heads radiate, in dense leafy cymes; involucre 5.5–7.5 mm, 3–4 mm diam, obconic; phyllaries 20–30 in 4–5 series, lanceolate, acute to acuminate, glabrous
Ray flowers 1–7; corollas 2–3 mm
Disk flowers 4–8; corollas 4.5–6.5 mm
Fruit 4.5–5.5 mm, cylindric, faintly 5-angled, glabrous to densely hairy; pappus = disk corollas, light brown
Chromosomes: 2n=18
Ecology: Rocky soils, cliffs
Elevation: 2100–2800 m.
Bioregional distribution: c East of Sierra Nevada, Desert Mountains
Distribution outside California: to Washington, Idaho, Utah
Flowering time: Jul–Nov
Synonyms: Haplopappus n. (Nutt.) D.C. Eaton