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POLYGONACEAE

BUCKWHEAT FAMILY

James C. Hickman

Annual to trees, some dioecious
Stem: nodes often swollen
Leaves simple, basal or cauline, alternate, opposite, or whorled, generally entire; stipules 0 or obvious and fused into a generally scarious sheath around stem
Inflorescence: small cluster, axillary or arrayed in cymes or panicles; involucres sometimes subtending 1–many flowers
Flower generally bisexual, small, ± radial; perianth generally 5–6-lobed, base ± tapered, often jointed to pedicel; stamens 2–9, often in 2 whorls; ovary superior, styles generally 3, generally fused at base
Fruit: achene, generally enclosed by persistent perianth, generally 3-angled, ovoid, and glabrous
Genera in family: 50 genera, 1100 species: worldwide, especially n temp; some cultivated for food (Fagopyrum ; Rheum , rhubarb; Rumex , sorrel) or ornamental (Antigonon , coral-vine; Muehlenbeckia ; Polygonum )
Reference: [Ronse Decraene & Akeroyd 1988 Bot J Linn Soc 98:321–371; Reveal et al. 1989 Phytologia 66(2–4):83–414]
Treatments of the 15 eriogonoid genera are based on the monographic work of James L. Reveal, who is gratefully acknowledged.

CHORIZANTHE

SPINEFLOWER

Annual, perennial herb, glabrous or hairy, sometimes glandular
Stem generally scapose (made up of inflorescence axes)
Leaves basal (rarely some cauline); stipule 0; blade generally ± oblanceolate
Inflorescence open or of few heads, sometimes 1-sided; bracts generally opposite, leaf-like to scale-like; involucres 1–several per axil, sessile, tube cylindric to bell-shaped, generally ± cross-ridged or net-veined, bracts (and ribs) 3–6, awns straight or hooked; flowers 1–2 per involucre
Flower: perianth white to red or yellow, lobes 6, entire to fringed or toothed; stamens 3–9
Fruit 1.5–4.5 mm, generally ± brown, glabrous
Species in genus: 50 species: temp w North America, sw South America
Etymology: (Greek: divided flower, from perianth)
Reference: [Reveal & Hardham 1989 Phytologia 66(2):98–198]

Native

C. breweri S. Watson

BREWER'S SPINEFLOWER


Stem decumbent to ascending, 3–35 cm, reddish, thinly hairy
Leaf: blade 5–20 mm, generally ovate, thinly hairy to tomentose below
Inflorescence: lower bracts ± oblanceolate, ± leaf-like, persistent, awns straight; involucres in ± open clusters, tube generally 2.5–3 mm, ± bell-shaped, hairs slender, curly, bracts 6, awns hooked
Flower: perianth 3–3.5 mm, white to red, hairy, lobes unequal, generally entire; stamens 9
Chromosomes: n=19
Ecology: Chaparral, foothill woodland, on serpentine
Elevation: < 800 m.
Bioregional distribution: Outer South Coast Ranges (sw San Luis Obispo Co.).Closely related to C. staticoides.

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