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POLEMONIACEAE

PHLOX FAMILY

Robert W. Patterson, Family Editor

Annual, perennial herb, shrub, vine
Leaves simple or compound, cauline (or most in basal rosette), alternate or opposite; stipules 0
Inflorescence: cymes, heads, or flowers solitary
Flower: calyx generally 5-ribbed, ribs often connected by translucent membranes that are generally torn by growing fruit; corolla generally 5-lobed, radial or bilateral, salverform to bell-shaped, throat often well defined; stamens generally 5, epipetalous, attached at same or different levels, filaments of same or different lengths, pollen white, yellow, blue, or red; ovary superior, chambers generally 3, style 1, stigmas generally 3
Fruit: capsule
Seeds 1–many, gelatinous or not when wet
Genera in family: 19 genera, 320 species: Am, n Eur, n Asia; some cultivated (Cantua, Cobaea (cup-and-saucer vine), Collomia, Gilia, Ipomopsis, Linanthus, Phlox )
Recent taxonomic note: *See also revised taxonomy of Porter and Johnson 2000 Aliso 19(1):55–91; Porter 1998 Aliso 17:83–85
.

GILIA

Alva G. Day

Annual, perennial herb, generally erect
Stem glabrous, hairy, glandular, or cobwebby
Leaves simple, generally alternate; basal generally in rosette, toothed, pinnately lobed, or entire; cauline generally reduced; leaf tips, calyx lobes acute, acuminate, or needle-like
Inflorescence: flowers solitary or clustered, 1–many in axils of bracts
Flower: calyx membranous between ribs, membrane splitting or expanding; corolla > calyx, lobes generally ovate
Fruit generally ovoid; chambers 3, valves separating from top
Seeds 3–many, brown, generally gelatinous when wet
Species in genus: ± 70 species: w North America, South America
Etymology: (Felipe Gil, 18th century Spanish botanist)

Native

G. brecciarum M.E. Jones

Plant sometimes with skunk-like odor
Stem 8–35 cm, densely cobwebby near base
Leaves: basal in semi-erect cluster, 2–5 cm, 2-pinnate; lobes > axis width; upper cauline lobes generally finger-like, middle lobe widest
Inflorescence: flowers (and generally fruits) clustered; pedicel glands dense, minute, black, stalked
Flower: calyx lobes acute, spreading, ribs thick, densely glandular, width > membrane, membrane generally purple-tinged
Fruit 4–7 mm, widely ovoid
Ecology: Open, sandy places
Elevation: 500–2300 m.
Bioregional distribution: c&s High Sierra Nevada, Tehachapi Mountain Area, Inner South Coast Ranges, Western Transverse Ranges, n edge San Gabriel Mountains, Great Basin Floristic Province, Mojave Desert
Distribution outside California: to se Oregon, Nevada, Utah
Sspp. intergrade somewhat in s CA.

Native

subsp. neglecta A.D. Grant & V.E. Grant


Stem spreading to erect
Flower: corolla showy, 10–20 mm, tube purple, generally exserted, throat widely V-shaped, lower half purple, white above, yellow spotted, lobes white, tips lavender; stamens, style long-exserted
Chromosomes: 2n=18
Ecology: Sandy desert flats
Elevation: 650–2100 m.
Bioregional distribution: c&s High Sierra Nevada, Tehachapi Mountain Area, Mojave Desert
Flowering time: Mar–May
Synonyms: subsp. argusana A.D. Grant & V.E. Grant
Horticultural information: TRY.

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