Jepson Flora Project: Jepson Interchange    

link to manual TREATMENT FROM THE JEPSON MANUAL (1993) previous taxon | next taxon
Jepson Interchange (more information)
©Copyright 1993 by the Regents of the University of California
For up-to-date information about California vascular plants, visit the Jepson eFlora.



Warren L. Wagner, except as specified Peter H. Raven, Family Coordinator

Annual to tree
Leaves basal or cauline, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous
Inflorescence: spike, raceme, panicle, or flowers solitary in axils; bracted
Flower generally bisexual, generally radial, opening at dawn or dusk; hypanthium sometimes prolonged beyond ovary (measured from ovary tip to sepal base); sepals generally 4(2–7); petals generally 4 (or as many as sepals, rarely 0), often "fading" darker; stamens generally 4 or 8(2), anthers 2-chambered, opening lengthwise, pollen generally interconnected by threads; ovary inferior, chambers generally 4 (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, or hemispheric
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like)
Seeds sometimes winged or hair-tufted
Genera in family: 15 genera, ± 650 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Gaura, Oenothera )
Reference: [Munz 1965 North America Fl II 5:1–278]



Annual, biennial, perennial herb, generally from taproot
Leaves basal or cauline, alternate, generally pinnately toothed to lobed
Inflorescence: spike, raceme-like, or flowers in axils of upper, reduced leaves
Flower radial, generally opening at dusk; sepals 4, reflexed in flower (sometimes 2–3 remaining adherent); petals 4, yellow, white, rose, or ± purple, generally fading orangish to purplish, tip notched or toothed; stamens 8, anthers attached at middle; ovary chambers 4, stigma deeply lobed, generally > anthers and cross-pollinated (or ± = anthers and self-pollinated)
Fruit cylindric to 4-winged, straight to curved, generally sessile (base sometimes seedless, stalk-like)
Seeds in generally 2(1–3) rows per chamber, or clustered
Species in genus: 119 species: Am, some widely naturalized
Etymology: (Greek: wine-scented)
Reference: [Dietrich & Wagner 1988 Syst Bot Monogr 24:1–91]
Many species self-pollinated; some of these have chromosome peculiarities (ring of 14 in meiosis) and ± 50% pollen fertility; they yield genetically ± identical offspring; they are identified as Permanent Translocation Heterozygote.


O. caespitosa Nutt.


Perennial, rosetted; caudex woody, new shoots generally from lateral roots; hairs glandular and sometimes also coarse and non-glandular
Stem sprawling, < 2 dm, or ± 0
Leaf 1.7–36 cm, oblanceolate to narrowly elliptic, generally irregularly dentate to lobed
Inflorescence: flowers in axils
Flower: hypanthium 30–165 mm; sepals 16–50 mm, tips in bud not free; petals 16–56 mm, white
Fruit 10–68 mm, 4–9 mm wide, cylindric to elliptic-ovate, tubercled
Seed obovate to ± triangular, papillate or netted, 1 side with a cavity sealed by a depressed, generally splitting membrane
Chromosomes: 2n=14,28
Ecology: Open desert scrub, pinyon/juniper woodland, coniferous and bristlecone-pine forests
Elevation: 1100–3400 m.
Bioregional distribution: Modoc Plateau (likely), East of Sierra Nevada, Desert
Distribution outside California: w US
Cross-pollinated. 5 intergrading subspp., 2 in CA.


subsp. crinita (Rydb.) Munz


Plant loosely to densely cespitose
Flower: hypanthium 30–85 mm; petals fading rose to purple
Fruit 10–34 mm, lanceolate to elliptic-ovate, generally S-shaped; stalk-like base 0–1 mm
Seed 2.9–3.5 mm; cavity margin lobed
Ecology: UNCOMMON. Calcium soils in bristlecone-pine forest, pinyon/juniper woodland, desert scrub
Elevation: 1150–3370 m.
Bioregional distribution: Modoc Plateau (likely), East of Sierra Nevada, Desert
Distribution outside California: w US
Flowering time: Jun–Sep
Synonyms: var. c. (Rydb.) Munz
2 intergrading forms differ in elevation, habit, leaf size, petal color; more study needed
Horticultural information: TRY.

previous taxon | next taxon
bioregional map for OENOTHERA%20caespitosa%20subsp.%20crinita being generated
YOU CAN HELP US make sure that our distributional information is correct and current. If you know that a plant occurs in a wild, reproducing state in a Jepson bioregion NOT highlighted on the map, please contact us with that information. Please realize that we cannot incorporate range extensions without access to a voucher specimen, which should (ultimately) be deposited in an herbarium. You can send the pressed, dried collection (with complete locality information indicated) to us (e-mail us for details) or refer us to an accessioned herbarium specimen. Non-occurrence of a plant in an indicated area is difficult to document, but we will especially value your input on those types of possible errors (see automatic conversion of distribution data to maps).
Overlay Consortium of California Herbaria specimen data by county on this map
Show other taxa with the same California distribution | Read about bioregions | Get lists of plants in a bioregion
Return to the Jepson Interchange main page
Return to treatment index page
  • This page is no longer being maintained.

University & Jepson Herbaria Home Page |
General Information | University Herbarium | Jepson Herbarium |
Visiting the Herbaria | On-line Resources | Research |
Education | Related Sites
Copyright © by the Regents of the University of California