Jepson Flora Project: Jepson Interchange    

 
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ONAGRACEAE

EVENING PRIMROSE FAMILY

Warren L. Wagner, except as specified Peter H. Raven, Family Coordinator

Annual to tree
Leaves basal or cauline, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous
Inflorescence: spike, raceme, panicle, or flowers solitary in axils; bracted
Flower generally bisexual, generally radial, opening at dawn or dusk; hypanthium sometimes prolonged beyond ovary (measured from ovary tip to sepal base); sepals generally 4(2–7); petals generally 4 (or as many as sepals, rarely 0), often "fading" darker; stamens generally 4 or 8(2), anthers 2-chambered, opening lengthwise, pollen generally interconnected by threads; ovary inferior, chambers generally 4 (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, or hemispheric
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like)
Seeds sometimes winged or hair-tufted
Genera in family: 15 genera, ± 650 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Gaura, Oenothera )
Reference: [Munz 1965 North America Fl II 5:1–278]

CAMISSONIA

SUN CUP

Annual, perennial herb, from taproot or lateral roots
Leaves basal, cauline, or both, alternate, simple to 2-pinnate
Inflorescence bracted; spike, raceme, or flowers solitary in axils
Flower radial, generally opening at dawn (rarely at dusk); sepals 4, reflexed (sometimes 2–3 remaining adherent); petals 4, yellow, white, lavender, often with darker basal spots, generally fading purplish or reddish; stamens (4)8, longer ones opposite sepals, anthers generally attached at middle (or base), pollen grains 3-angled except in polyploid taxa (visible with hand lens); ovary chambers 4, stigma ± head-like or hemispheric, generally > anthers and cross-pollinated (or ± = anthers and self-pollinated)
Fruit straight to coiled, generally sessile
Seeds in 1–2 rows per chamber
Species in genus: 62 species: w North America (especially CA-FP), 1 South America
Etymology: (L.A. von Chamisso, French-born German botanist, 1781–1838)
Reference: [Raven 1969 Contr US Natl Herb 37:161–396]
Polyploidy and self-pollination have predominated in evolution of genus. Previously included in Oenothera (" O." in synonyms).

Native

C. ignota (Jeps.) P.H. Raven

Annual, rosetted, generally reddish, minutely strigose (inflorescence glabrous, longer-hairy, or glandular)
Stem ascending or erect, < 55 cm, ± fleshy
Leaf < 60 mm, narrowly elliptic to narrowly ovate, minutely serrate; petiole < 25 mm
Inflorescence nodding
Flower: hypanthium 1.1–3 mm; sepals 2.5–5.5 mm; petals (3)4.8–8 mm, yellow fading reddish, bases red-dotted or not; < 5% of pollen grains 4–5-angled
Fruit generally 20–30 mm, 0.8–1 mm wide, cylindric (drying 4-angled), < 5-coiled
Seeds in 1 row per chamber, ± 1.2 mm, minutely pitted in rows, dull brownish black
Chromosomes: 2n=14
Ecology: Generally clay fields, slopes in coastal-sage scrub or chaparral, sandy soils in mtns
Elevation: 100–1100 m.
Bioregional distribution: c High Sierra Nevada (Madera Co.), Sacramento Valley (uncommon), Central Western California, Southwestern California (including Santa Cruz Island)
Distribution outside California: n Baja California
Synonyms: O. i. (Jeps.) Munz
Self-pollinated.

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