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  • Up-to-date information about California vascular plants is available from the Jepson eFlora.

ONAGRACEAE

EVENING PRIMROSE FAMILY

Warren L. Wagner, except as specified Peter H. Raven, Family Coordinator

Annual to tree
Leaves basal or cauline, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous
Inflorescence: spike, raceme, panicle, or flowers solitary in axils; bracted
Flower generally bisexual, generally radial, opening at dawn or dusk; hypanthium sometimes prolonged beyond ovary (measured from ovary tip to sepal base); sepals generally 4(2–7); petals generally 4 (or as many as sepals, rarely 0), often "fading" darker; stamens generally 4 or 8(2), anthers 2-chambered, opening lengthwise, pollen generally interconnected by threads; ovary inferior, chambers generally 4 (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, or hemispheric
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like)
Seeds sometimes winged or hair-tufted
Genera in family: 15 genera, ± 650 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Gaura, Oenothera )
Reference: [Munz 1965 North America Fl II 5:1–278]

CAMISSONIA

SUN CUP

Annual, perennial herb, from taproot or lateral roots
Leaves basal, cauline, or both, alternate, simple to 2-pinnate
Inflorescence bracted; spike, raceme, or flowers solitary in axils
Flower radial, generally opening at dawn (rarely at dusk); sepals 4, reflexed (sometimes 2–3 remaining adherent); petals 4, yellow, white, lavender, often with darker basal spots, generally fading purplish or reddish; stamens (4)8, longer ones opposite sepals, anthers generally attached at middle (or base), pollen grains 3-angled except in polyploid taxa (visible with hand lens); ovary chambers 4, stigma ± head-like or hemispheric, generally > anthers and cross-pollinated (or ± = anthers and self-pollinated)
Fruit straight to coiled, generally sessile
Seeds in 1–2 rows per chamber
Species in genus: 62 species: w North America (especially CA-FP), 1 South America
Etymology: (L.A. von Chamisso, French-born German botanist, 1781–1838)
Reference: [Raven 1969 Contr US Natl Herb 37:161–396]
Polyploidy and self-pollination have predominated in evolution of genus. Previously included in Oenothera (" O." in synonyms).

Native

C. cheiranthifolia (Spreng.) Raim.

BEACH EVENING PRIMROSE

Perennial or subshrub, short-lived, rosetted, densely strigose (rarely glabrous); hairs of inflorescence generally erect, short
Stems prostrate to ± ascending, < 60(130) cm, peeling
Leaf 5–50 mm, narrowly ovate to obovate, minutely serrate; cauline petioles 0–10 mm
Inflorescence nodding
Flower: hypanthium 2.1–8.5 mm; sepals 4–11.5 mm; petals 6–20 mm, yellow fading reddish, bases with 0–2 red dots
Fruit 10–25 mm, 2–2.5 mm wide, 4-angled, generally 1–2-coiled
Seeds in 1 row per chamber, 1.2–1.3 mm, minutely pitted in rows, dull brownish black
Chromosomes: 2n=14
Ecology: Sandy slopes, flats, coastal dunes
Elevation: < 100 m.
Bioregional distribution: North Coast, Central Coast, South Coast, Channel Islands
Distribution outside California: sw Oregon, n Baja California
Synonyms: O. c. Spreng. including var. nitida (Greene) Munz
Generally cross-pollinated. Sspp. intergrade on ChI.

Native

subsp. suffruticosa (S. Watson) P.H. Raven

Subshrub; hairs generally dense and silvery
Flower: hypanthium 5–8.5 mm; petals (10)12–20 mm, bases with 1–2 red dots; anthers 2.2–3 mm; style 13–23 mm
Ecology: Habitats of sp.
Bioregional distribution: South Coast
Distribution outside California: n Baja California
Synonyms: O. c. subsp. s. (S. Watson) Munz
Generally cross-pollinated (± self-incompatible). Hybridizes widely with C. bistorta.
Horticultural information: SUN: 15, 16, 17, 24 &IRR: 14, 19, 20, 21, 22, 23.

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