TREATMENT FROM THE JEPSON MANUAL	previous taxon | next taxon Jepson Interchange (more information)
	©Copyright 1993 by the Regents of the University of California Print edition is available from the University of California Press

	The second edition of The Jepson Manual (2012) is available from the University of California Press
	See also the Jepson eFlora, which parallels the Second Edition

ONAGRACEAE

EVENING PRIMROSE FAMILY

Warren L. Wagner, except as specified Peter H. Raven, Family Coordinator

Annual to tree
Leaves basal or cauline, alternate, opposite, or whorled, generally simple and toothed (to pinnately compound); stipules 0 or generally deciduous
Inflorescence: spike, raceme, panicle, or flowers solitary in axils; bracted
Flower generally bisexual, generally radial, opening at dawn or dusk; hypanthium sometimes prolonged beyond ovary (measured from ovary tip to sepal base); sepals generally 4(2–7); petals generally 4 (or as many as sepals, rarely 0), often "fading" darker; stamens generally 4 or 8(2), anthers 2-chambered, opening lengthwise, pollen generally interconnected by threads; ovary inferior, chambers generally 4 (sometimes becoming 1), placentas axile or parietal, ovules 1–many per chamber, style 1, stigma 4-lobed (or lobes as many as sepals), club-shaped, or hemispheric
Fruit: capsule, loculicidal (sometimes berry or indehiscent and nut-like)
Seeds sometimes winged or hair-tufted
Genera in family: 15 genera, ± 650 species: worldwide, especially w North America; many cultivated (Clarkia, Epilobium, Fuchsia, Gaura, Oenothera )
Reference: [Munz 1965 North America Fl II 5:1–278]

CAMISSONIA

SUN CUP

Annual, perennial herb, from taproot or lateral roots
Leaves basal, cauline, or both, alternate, simple to 2-pinnate
Inflorescence bracted; spike, raceme, or flowers solitary in axils
Flower radial, generally opening at dawn (rarely at dusk); sepals 4, reflexed (sometimes 2–3 remaining adherent); petals 4, yellow, white, lavender, often with darker basal spots, generally fading purplish or reddish; stamens (4)8, longer ones opposite sepals, anthers generally attached at middle (or base), pollen grains 3-angled except in polyploid taxa (visible with hand lens); ovary chambers 4, stigma ± head-like or hemispheric, generally > anthers and cross-pollinated (or ± = anthers and self-pollinated)
Fruit straight to coiled, generally sessile
Seeds in 1–2 rows per chamber
Species in genus: 62 species: w North America (especially CA-FP), 1 South America
Etymology: (L.A. von Chamisso, French-born German botanist, 1781–1838)
Reference: [Raven 1969 Contr US Natl Herb 37:161–396]
Polyploidy and self-pollination have predominated in evolution of genus. Previously included in Oenothera (" O." in synonyms).

Native

C. californica (Torr. & A. Gray) P.H. Raven

Annual, subglabrous (or leaves minutely strigose, inflorescence sparsely glandular-hairy); rosette well developed
Stem straight, slender, erect, 2–180 cm, ± glaucous or green
Leaves much reduced upward, < 300 mm, narrowly elliptic, irregularly and sharply pinnately lobed
Inflorescence: flowers widely spaced
Flower: hypanthium 0.6–1.5 mm, closed by fleshy disk; sepals 3.9–8 mm; petals 6–14 mm, yellow, base ± red-flecked
Fruit reflexed, 45–110 mm, cylindric (drying 4-angled), ± straight
Seeds in 1 row per chamber, 1.3–1.6 mm, olive with purple spots
Chromosomes: 2n=14, 28
Ecology: Open places in coastal-sage scrub, chaparral, desert scrub
Elevation: < 1300 m.
Bioregional distribution: s Outer North Coast Ranges (Sonoma Co.), San Joaquin Valley (Fresno Co.), South Coast Ranges, Southwestern California, Desert
Distribution outside California: sw Arizona, nw Mexico
Flowering time: Apr–Jun
Synonyms: Eulobus c. Torr. & A. Gray; O. leptocarpa Greene
Self-pollinated
Horticultural information: SUN, DRY, DRN: 7, 8, 9, 10, 11, 12, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24.